# Stream Consumers and Lotic Ecosystem Rates (SCALER) in streams of the HJ Andrews Experimental Forest, 2014-2016

## Dataset

- **Database Code:** AN008

- **Title:** Stream Consumers and Lotic Ecosystem Rates (SCALER) in streams of the HJ Andrews Experimental Forest, 2014-2016

- **Abstract:**

SCALER attempts to address the question of whether small-scale ecological experiments can be applied to understand the behavior of entire ecological systems. Specifically, the role of comsumers to stream functional processes were examined by measurements of structure and function within stream reaches. These results feed into a broader continent wide examination within a variety of ecological systems. Rates of stream metabolism (photosynthesis and respiration) and nutrient uptake were measured, as well as the way these rates respond to animal exclusions (used to mimic loss of animal diversity in streams). Aquatic food web and biogeochemical responses in three consecutive 75m stream reaches were measured by manipulating densities of fish and amphibians (natural, depletion, and addition). Additionally, various stream characteristics were measured including streamflow, water chemistry, nutrients, metabolism, stream habitat measurements (wetted widths, depths and substrate), and canopy conditions (percent shade). The experiment was carried out every summer between 2014-2016 in streams of the HJ Andrews Experimental Forest to capture interannual variability.


- **Keywords:** stream ecology; biomass; species abundance; respiration; metabolism; primary production; nutrients; streams; invertebrates; amphibians; fishes; primary production

- **Temporal coverage:** 2014-08-14 to 2016-08-17

- **Principal Investigator:** Brooke E. Penaluna


## Data Entities

1. AN00801
2. AN00802
3. AN00803
4. AN00804
5. AN00805
6. AN00806
7. AN00807
8. AN00808

## Entity 1: AN00801

### Attribute list

- **DBCODE** — varchar, enum

FSDB Database Code

  - **AN008** — FSDB Database Study Code


- **ENTITY** — numeric(1), range  1.0000  1.0000, number

Entity number


- **SAMPLE_TYPE** — varchar, enum

Spaitial scale of sampling

  - **REACH** — Sampling conducted at reach scale, typically 50 to 65m stretch
  - **PATCH** — Sampling conducted at patch scale in plots or chamber units within reach


- **YEAR** — numeric(1), range  2014.0000  2016.0000, year (yyyy)

Year sample collected


- **SITE_NUMBER** — varchar, enum

ID number assigned to site

  - **SCALER_1** — SCALER sample site 1 on McRae Creek
  - **SCALER_2** — SCALER sample site 2, on McRae Creek at the STREON site
  - **SCALER_3** — SCALER sample site 3 on Lookout Creek


- **REACH_TREATMENT** — varchar, enum

Reach treatment

  - **C** — Control reach (consumers ambient)
  - **D** — Treatment/deletion reach (consumers removed)
  - **A** — Addition reach (consumers added)


- **SAMPLE_REPLICATE** — numeric(1), range  1.0000  7.0000, number

Sample replicate number


- **SAMPLE_TIMING** — varchar, enum

Type of sample timing collection during season

  - **START** — Beginning of season first sampling
  - **END** — End of season second and final sampling


- **SAMPLING_DATE** — datetime, freetext

Date of sample collection


- **EPILITHON_BIOMASS** — numeric(1), range  0.8800  52.0800, grams per square meter

Biomass (AFDM) concentration


- **CHL_A** — numeric(1), range  0.9500  35.4800, milligrams per square meter

Chlorophyll a concentration corrected following Parker et al. (2016)


- **NOTES** — varchar, freetext

Notes taken during sample collection and analysis


## Entity 2: AN00802

### Attribute list

- **DBCODE** — varchar, enum

FSDB Database Code

  - **AN008** — FSDB Database Study Code


- **ENTITY** — numeric(1), range  2.0000  2.0000, number

Entity number


- **YEAR** — numeric(1), range  2014.0000  2016.0000, year (yyyy)

Year sample collected


- **SITE_NUMBER** — varchar, enum

ID number assigned to site

  - **SCALER_1** — SCALER sample site 1 on McRae Creek
  - **SCALER_2** — SCALER sample site 2, on McRae Creek at the STREON site
  - **SCALER_3** — SCALER sample site 3 on Lookout Creek


- **REACH_TREATMENT** — varchar, enum

Reach treatment

  - **C** — Control reach (consumers ambient)
  - **D** — Treatment/deletion reach (consumers removed)
  - **A** — Addition reach (consumers added)


- **SAMPLING_DATE** — datetime, freetext

Date of sample collection


- **DAY_OF_EXPERIMENT** — numeric(1), range  1.0000  50.0000, number

Day of experiment (Number since start)


- **PASS** — numeric(1), range  1.0000  11.0000, number

Pass through reach when collected (multiple passes made during collection)


- **DIRECTION** — varchar, enum

Direction headed while sampling (upstream or downstream)

  - **U** — Direction of pass through stream to collect consumers is upstream
  - **D** — Direction of pass through stream to collect consumers is downstream


- **SPECIES** — varchar, enum

Consumer species code

  - **PS** — Cottus beldingi
  - **CT** — Oncorhyncus clarkii clarkii
  - **TR** — Oncorhyncus mykiss
  - **LD** — Rhinichthys cataractae
  - **TF** — Ascaphus truei
  - **GS** — Dicamptodon tenebrosus
  - **DU** — Plethodon dunni
  - **RF** — Rana aurora
  - **CF** — Rana cascadae
  - **RSN** — Taricha granulosa


- **VERT_INDEX** — numeric(1), range  1.0000  172.0000, number

unique index of site, reach, date, pass, direction, species sampled


- **LENGTH2_AMPHIBS** — numeric(1), range  1.0000  200.0000, millimeters

VL; vent length, amphibs only


- **LENGTH** — numeric(1), range  1.0000  300.0000, millimeters

Length of consumer


- **WEIGHT** — numeric(1), range  0.1000  200.0000, grams

Weight of consumer


- **FROG_DEV_STAGE** — varchar, enum

Stage of frog development

  - **NA** — Not applicable, data either misssing or does not apply to species
  - **A** — Adult
  - **FHL** — Full front and hind legs, tail remaining
  - **HL** — Hind legs fully developed, front legs undeveloped
  - **NL** — No legs, tadpole stage
  - **SHL** — Stubby hind legs, front legs undeveloped
  - **SFL** — Stubby front legs, hind legs developed


- **NOTES** — varchar, freetext

Notes taken during sample collection and analysis


- **QA_NOTES** — varchar, freetext

Note recorded during QA/QC


## Entity 3: AN00803

### Attribute list

- **DBCODE** — varchar, enum

FSDB Database Code

  - **AN008** — FSDB Database Study Code


- **ENTITY** — numeric(1), range  3.0000  3.0000, number

Entity number


- **YEAR** — numeric(1), range  2014.0000  2016.0000, year (yyyy)

Year sample collected


- **SITE_NUMBER** — varchar, enum

ID number assigned to site

  - **SCALER_1** — SCALER sample site 1 on McRae Creek
  - **SCALER_2** — SCALER sample site 2, on McRae Creek at the STREON site
  - **SCALER_3** — SCALER sample site 3 on Lookout Creek


- **REACH_TREATMENT** — varchar, enum

Reach treatment

  - **C** — Control reach (consumers ambient)
  - **D** — Treatment/deletion reach (consumers removed)
  - **A** — Addition reach (consumers added)


- **SAMPLING_DATE** — datetime, freetext

Date of sample collection


- **SAMPLE_TIMING** — varchar, enum

Type of sample timing collection during season

  - **START** — Beginning of season first sampling
  - **END** — End of season second and final sampling


- **UTN** — numeric(1), range  0.0010  1.0000, milligrams per liter

Total unfiltered nitrogen (N) concentration


- **NH4_N** — numeric(1), range  0.0010  1.0000, milligrams per liter

Ammonium (NH4-N) concentration


- **NO3_N** — numeric(1), range  0.0010  1.0000, milligrams per liter

Nitrate (NO3-N) concentration


- **UTP** — numeric(1), range  0.0010  1.0000, milligrams per liter

Total unfiltered phosphorous (P) concentration


- **PO4_P** — numeric(1), range  0.0010  1.0000, milligrams per liter

Orthophosphate (PO4-P) concentration


- **DOC** — numeric(1), range  0.0010  1.0000, milligrams per liter

Dissolved Organic Carbon (mg C/L) concentration


## Entity 4: AN00804

### Attribute list

- **DBCODE** — varchar, enum

FSDB Database Code

  - **AN008** — FSDB Database Study Code


- **ENTITY** — numeric(1), range  4.0000  4.0000, number

Entity number


- **YEAR** — numeric(1), range  2014.0000  2016.0000, year (yyyy)

Year sample collected


- **SITE_NUMBER** — varchar, enum

ID number assigned to site

  - **SCALER_1** — SCALER sample site 1 on McRae Creek
  - **SCALER_2** — SCALER sample site 2, on McRae Creek at the STREON site
  - **SCALER_3** — SCALER sample site 3 on Lookout Creek


- **REACH_TREATMENT** — varchar, enum

Reach treatment

  - **C** — Control reach (consumers ambient)
  - **D** — Treatment/deletion reach (consumers removed)
  - **A** — Addition reach (consumers added)


- **SAMPLING_DATE** — datetime, freetext

Date of sample collection


- **SAMPLE_TIMING** — varchar, enum

Type of sample timing collection during season

  - **START** — Beginning of season first sampling
  - **END** — End of season second and final sampling


- **TRANSECT_M** — varchar, enum

Number of meters from bottom of reach where measurements were taken

  - **** — Most upstream transect in study reach
  - **5** — Second transect, 5 meters downstream from transect 0
  - **10** — Third transect, 10 m downstream of transect 0
  - **15** — Fourth transect, 15 m downstream of transect 0
  - **20** — Fifth transect, 20 m downstream of transect 0
  - **25** — Sixth transect, 25 m downstream of transect 0
  - **30** — Seventh transect, 30 m downstream of transect 0
  - **35** — Eighth transect, 35 m downstream of transect 0
  - **40** — Ninth transect, 40 m downstream of transect 0
  - **45** — Tenth transect, 45 m downstream of transect 0
  - **50** — Eleventh transect, 50 m downstream of transect 0
  - **55** — Twelfth transect, 55 m downstream of transect 0
  - **60** — Thirteenth transect, 60 m downstream of transect 0
  - **65** — Fourteenth transect, 65 m downstream of transect 0
  - **70** — Fifteenth transect, 70 m downstream of transect 0
  - **75** — Sixteenth transect, 75 m downstream of transect 0


- **AVERAGE_WIDTH_M** — numeric(1), range  1.8000  11.0000, meters

Wetted width measured at cross section of reach


## Entity 5: AN00805

### Attribute list

- **DBCODE** — varchar, enum

FSDB Database Code

  - **AN008** — FSDB Database Study Code


- **ENTITY** — numeric(1), range  5.0000  5.0000, number

Entity number


- **YEAR** — numeric(1), range  2014.0000  2016.0000, year (yyyy)

Year sample collected


- **SITE_NUMBER** — varchar, enum

ID number assigned to site

  - **SCALER_1** — SCALER sample site 1 on McRae Creek
  - **SCALER_2** — SCALER sample site 2, on McRae Creek at the STREON site
  - **SCALER_3** — SCALER sample site 3 on Lookout Creek


- **REACH_TREATMENT** — varchar, enum

Reach treatment

  - **C** — Control reach (consumers ambient)
  - **D** — Treatment/deletion reach (consumers removed)
  - **A** — Addition reach (consumers added)


- **SAMPLING_DATE** — datetime, freetext

Date of sample collection


- **TRANSECT_M** — varchar, enum

Number of meters from bottom of reach where measurements were taken

  - **** — Most upstream transect in study reach
  - **5** — Second transect, 5 meters downstream from transect 0
  - **10** — Third transect, 10 m downstream of transect 0
  - **15** — Fourth transect, 15 m downstream of transect 0
  - **20** — Fifth transect, 20 m downstream of transect 0
  - **25** — Sixth transect, 25 m downstream of transect 0
  - **30** — Seventh transect, 30 m downstream of transect 0
  - **35** — Eighth transect, 35 m downstream of transect 0
  - **40** — Ninth transect, 40 m downstream of transect 0
  - **45** — Tenth transect, 45 m downstream of transect 0
  - **50** — Eleventh transect, 50 m downstream of transect 0
  - **55** — Twelfth transect, 55 m downstream of transect 0
  - **60** — Thirteenth transect, 60 m downstream of transect 0
  - **65** — Fourteenth transect, 65 m downstream of transect 0
  - **70** — Fifteenth transect, 70 m downstream of transect 0
  - **75** — Sixteenth transect, 75 m downstream of transect 0


- **SAMPLE_PT** — numeric(1), range  1.0000  56.0000, number

Sampling point across the transect


- **SUBSTRATE_MM** — numeric(1), range  0.1000  2000.0000, millimeters

Width of substrate


- **DEPTH_CM** — numeric(1), range  0.0000  1000.0000, centimeters

Depth of stream at cross section


- **NOTES** — varchar, freetext

Notes taken during sample collection and analysis


## Entity 6: AN00806

### Attribute list

- **DBCODE** — varchar, enum

FSDB Database Code

  - **AN008** — FSDB Database Study Code


- **ENTITY** — numeric(1), range  6.0000  6.0000, number

Entity number


- **YEAR** — numeric(1), range  2014.0000  2016.0000, year (yyyy)

Year sample collected


- **SITE_NUMBER** — varchar, enum

ID number assigned to site

  - **SCALER_1** — SCALER sample site 1 on McRae Creek
  - **SCALER_2** — SCALER sample site 2, on McRae Creek at the STREON site
  - **SCALER_3** — SCALER sample site 3 on Lookout Creek


- **REACH_TREATMENT** — varchar, enum

Reach treatment

  - **C** — Control reach (consumers ambient)
  - **D** — Treatment/deletion reach (consumers removed)
  - **A** — Addition reach (consumers added)


- **SAMPLING_DATE** — datetime, freetext

Date of sample collection


- **TRANSECT_M** — varchar, enum

Number of meters from bottom of reach where measurements were taken

  - **** — Most upstream transect in study reach
  - **5** — Second transect, 5 meters downstream from transect 0
  - **10** — Third transect, 10 m downstream of transect 0
  - **15** — Fourth transect, 15 m downstream of transect 0
  - **20** — Fifth transect, 20 m downstream of transect 0
  - **25** — Sixth transect, 25 m downstream of transect 0
  - **30** — Seventh transect, 30 m downstream of transect 0
  - **35** — Eighth transect, 35 m downstream of transect 0
  - **40** — Ninth transect, 40 m downstream of transect 0
  - **45** — Tenth transect, 45 m downstream of transect 0
  - **50** — Eleventh transect, 50 m downstream of transect 0
  - **55** — Twelfth transect, 55 m downstream of transect 0
  - **60** — Thirteenth transect, 60 m downstream of transect 0
  - **65** — Fourteenth transect, 65 m downstream of transect 0
  - **70** — Fifteenth transect, 70 m downstream of transect 0
  - **75** — Sixteenth transect, 75 m downstream of transect 0


- **UP_PERCENT_UNSHADED** — numeric(1), range  0.0000  100.0000, percent

Percent of unshaded canopy looking upstream


- **LEFT_PERCENT_UNSHADED** — numeric(1), range  0.0000  100.0000, percent

Percent of unshaded canopy looking to left bank


- **DOWN_PERCENT_UNSHADED** — numeric(1), range  0.0000  100.0000, percent

Percent of unshaded canopy looking to right bank


- **RIGHT_PERCENT_UNSHADED** — numeric(1), range  0.0000  100.0000, percent

Percent of unshaded canopy looking downstream


## Entity 7: AN00807

### Attribute list

- **DBCODE** — varchar, enum

FSDB Database Code

  - **AN008** — FSDB Database Study Code


- **ENTITY** — numeric(1), range  7.0000  7.0000, number

Entity number


- **YEAR** — numeric(1), range  2014.0000  2016.0000, year (yyyy)

Year sample collected


- **SITE_NUMBER** — varchar, enum

ID number assigned to site

  - **SCALER_1** — SCALER sample site 1 on McRae Creek
  - **SCALER_2** — SCALER sample site 2, on McRae Creek at the STREON site
  - **SCALER_3** — SCALER sample site 3 on Lookout Creek


- **REACH_TREATMENT** — varchar, enum

Reach treatment

  - **C** — Control reach (consumers ambient)
  - **D** — Treatment/deletion reach (consumers removed)
  - **A** — Addition reach (consumers added)


- **SAMPLING_DATE** — datetime, freetext

Date of sample collection


- **SAMPLE_TIMING** — varchar, enum

Type of sample timing collection during season

  - **START** — Beginning of season first sampling
  - **END** — End of season second and final sampling


- **SAMPLE_REPLICATE** — numeric(1), range  1.0000  7.0000, number

Sample replicate number


- **FBOM_AFDM** — numeric(1), range  1.0000  400.0000, grams per square meter

Concentration of Fine Benthic Organic Matter (Ash Free Dry Mass)


- **NOTES** — varchar, freetext

Notes taken during sample collection and analysis


## Entity 8: AN00808

### Attribute list

- **DBCODE** — varchar, enum

FSDB Database Code

  - **AN008** — FSDB Database Study Code


- **ENTITY** — numeric(1), range  8.0000  8.0000, number

Entity number


- **YEAR** — numeric(1), range  2014.0000  2016.0000, year (yyyy)

Year sample collected


- **SITE_NUMBER** — varchar, enum

ID number assigned to site

  - **SCALER_1** — SCALER sample site 1 on McRae Creek
  - **SCALER_2** — SCALER sample site 2, on McRae Creek at the STREON site
  - **SCALER_3** — SCALER sample site 3 on Lookout Creek


- **REACH_TREATMENT** — varchar, enum

Reach treatment

  - **C** — Control reach (consumers ambient)
  - **D** — Treatment/deletion reach (consumers removed)
  - **A** — Addition reach (consumers added)


- **SAMPLE_LOCATION** — varchar, enum

Location sampled in reach

  - **BOTTOM** — Bottom or downstream ending point of sampling
  - **TOP** — Top or upstream starting point of sampling


- **SAMPLE_TIMING** — varchar, enum

Type of sample timing collection during season

  - **START** — Beginning of season first sampling
  - **END** — End of season second and final sampling


- **TIMESTAMP** — datetime, freetext

Date and time of sample collection (5 min samples)


- **SAMPLE_ID** — numeric(1), range  1.0000  52476.0000, number

Unique ID number assigned to sample


- **TEMPERATURE** — numeric(1), range  8.1800  17.6600, degrees Celsius

Temperature


- **PERCENT_DO** — numeric(1), range  80.0000  100.1000, percent

Percentage of dissolved oxygen


- **DO_MG_L** — numeric(1), range  8.0000  10.5000, milligrams per liter

Concentration of dissolved oxygen


- **EC_US_CM** — numeric(1), range  34.0000  2038.0000, micro siemens per centimeter

Electrical conductivity


## Attribute Definitions

**AVERAGE_WIDTH_M**

Wetted width measured at cross section of reach


**CHL_A**

Chlorophyll a concentration corrected following Parker et al. (2016)


**DAY_OF_EXPERIMENT**

Day of experiment (Number since start)


**DBCODE**

FSDB Database Code


**DEPTH_CM**

Depth of stream at cross section


**DIRECTION**

Direction headed while sampling (upstream or downstream)


**DOC**

Dissolved Organic Carbon (mg C/L) concentration


**DOWN_PERCENT_UNSHADED**

Percent of unshaded canopy looking to right bank


**DO_MG_L**

Concentration of dissolved oxygen


**EC_US_CM**

Electrical conductivity


**ENTITY**

Entity number


**EPILITHON_BIOMASS**

Biomass (AFDM) concentration


**FBOM_AFDM**

Concentration of Fine Benthic Organic Matter (Ash Free Dry Mass)


**FROG_DEV_STAGE**

Stage of frog development


**LEFT_PERCENT_UNSHADED**

Percent of unshaded canopy looking to left bank


**LENGTH**

Length of consumer


**LENGTH2_AMPHIBS**

VL; vent length, amphibs only


**NH4_N**

Ammonium (NH4-N) concentration


**NO3_N**

Nitrate (NO3-N) concentration


**NOTES**

Notes taken during sample collection and analysis


**PASS**

Pass through reach when collected (multiple passes made during collection)


**PERCENT_DO**

Percentage of dissolved oxygen


**PO4_P**

Orthophosphate (PO4-P) concentration


**QA_NOTES**

Note recorded during QA/QC


**REACH_TREATMENT**

Reach treatment


**RIGHT_PERCENT_UNSHADED**

Percent of unshaded canopy looking downstream


**SAMPLE_ID**

Unique ID number assigned to sample


**SAMPLE_LOCATION**

Location sampled in reach


**SAMPLE_PT**

Sampling point across the transect


**SAMPLE_REPLICATE**

Sample replicate number


**SAMPLE_TIMING**

Type of sample timing collection during season


**SAMPLE_TYPE**

Spaitial scale of sampling


**SAMPLING_DATE**

Date of sample collection


**SITE_NUMBER**

ID number assigned to site


**SPECIES**

Consumer species code


**SUBSTRATE_MM**

Width of substrate


**TEMPERATURE**

Temperature


**TIMESTAMP**

Date and time of sample collection (5 min samples)


**TRANSECT_M**

Number of meters from bottom of reach where measurements were taken


**UP_PERCENT_UNSHADED**

Percent of unshaded canopy looking upstream


**UTN**

Total unfiltered nitrogen (N) concentration


**UTP**

Total unfiltered phosphorous (P) concentration


**VERT_INDEX**

unique index of site, reach, date, pass, direction, species sampled


**WEIGHT**

Weight of consumer


**YEAR**

Year sample collected


## Enumerated Domains

### DBCODE

- **AN008** — FSDB Database Study Code

### SAMPLE_TYPE

- **REACH** — Sampling conducted at reach scale, typically 50 to 65m stretch

- **PATCH** — Sampling conducted at patch scale in plots or chamber units within reach

### SITE_NUMBER

- **SCALER_1** — SCALER sample site 1 on McRae Creek

- **SCALER_2** — SCALER sample site 2, on McRae Creek at the STREON site

- **SCALER_3** — SCALER sample site 3 on Lookout Creek

### REACH_TREATMENT

- **C** — Control reach (consumers ambient)

- **D** — Treatment/deletion reach (consumers removed)

- **A** — Addition reach (consumers added)

### SAMPLE_TIMING

- **START** — Beginning of season first sampling

- **END** — End of season second and final sampling

### DIRECTION

- **U** — Direction of pass through stream to collect consumers is upstream

- **D** — Direction of pass through stream to collect consumers is downstream

### SPECIES

- **PS** — Cottus beldingi

- **CT** — Oncorhyncus clarkii clarkii

- **TR** — Oncorhyncus mykiss

- **LD** — Rhinichthys cataractae

- **TF** — Ascaphus truei

- **GS** — Dicamptodon tenebrosus

- **DU** — Plethodon dunni

- **RF** — Rana aurora

- **CF** — Rana cascadae

- **RSN** — Taricha granulosa

### FROG_DEV_STAGE

- **NA** — Not applicable, data either misssing or does not apply to species

- **A** — Adult

- **FHL** — Full front and hind legs, tail remaining

- **HL** — Hind legs fully developed, front legs undeveloped

- **NL** — No legs, tadpole stage

- **SHL** — Stubby hind legs, front legs undeveloped

- **SFL** — Stubby front legs, hind legs developed

### TRANSECT_M

- **** — Most upstream transect in study reach

- **5** — Second transect, 5 meters downstream from transect 0

- **10** — Third transect, 10 m downstream of transect 0

- **15** — Fourth transect, 15 m downstream of transect 0

- **20** — Fifth transect, 20 m downstream of transect 0

- **25** — Sixth transect, 25 m downstream of transect 0

- **30** — Seventh transect, 30 m downstream of transect 0

- **35** — Eighth transect, 35 m downstream of transect 0

- **40** — Ninth transect, 40 m downstream of transect 0

- **45** — Tenth transect, 45 m downstream of transect 0

- **50** — Eleventh transect, 50 m downstream of transect 0

- **55** — Twelfth transect, 55 m downstream of transect 0

- **60** — Thirteenth transect, 60 m downstream of transect 0

- **65** — Fourteenth transect, 65 m downstream of transect 0

- **70** — Fifteenth transect, 70 m downstream of transect 0

- **75** — Sixteenth transect, 75 m downstream of transect 0

### SAMPLE_LOCATION

- **BOTTOM** — Bottom or downstream ending point of sampling

- **TOP** — Top or upstream starting point of sampling
