# Population dynamics of young forest stands as affected by density and nutrient regime in the Andrews Experimental Forest, 1981-1997 ## Dataset - **Database Code:** TP088 - **Title:** Population dynamics of young forest stands as affected by density and nutrient regime in the Andrews Experimental Forest, 1981-1997 - **Abstract:** To test hypotheses concerning feedback mechamisms (competition and mortality) which effect the dynamics of plant population development. Specifically, to test if manipulation of site quality (through fertilization) or the variance in individual tree leaf area (through pruning) will produce changes in the dominance hierarchy and thus in the growth mode of stands as they approach self-thinning (mortality). - **Keywords:** communities; populations; plant ecology; population dynamics; stand density; succession; primary production; growth; fertilization; fertilizer; forests; plants; Long-Term Ecological Research (LTER); populations; primary production - **Temporal coverage:** 1981-01-01 to 1997-01-01 - **Principal Investigator:** David A. Perry ## Data Entities 1. TP08801 2. TP08802 3. TP08803 4. TP08804 5. TP08805 6. TP08806 7. TP08807 ## Entity 1: TP08801 ### Attribute list - **DATACODE** — varchar, freetext Data set code - **FORMAT** — numeric(1), range 1.0000 1.0000, number Format number - **SITE** — varchar, enum Site identification code - **L701** — UNIT L701 - **L107** — UNIT L107 - **L111** — UNIT L111 - **L405** — UNIT L405 - **DENSITY** — varchar, enum Stocking density level - **C** — Control;no thinning within the stand - **M** — Medium;density was reduced to 25% of maximum stand density index - **L** — Low;density was reduced to 15% of maximum stand density index - **TMT** — varchar, enum Type of treatment applied to plots within stands - **F** — Fertilized - **PF** — Fertilized and pruned - **C** — Control - **P** — Pruned - **YEAR** — numeric(1), range 1981.0000 1981.0000, year (yyyy) Year of data collection - **PLOT15TR** — numeric(1), range 4.0000 34.0000, number Number of trees in a density plot centered at a random distance and direction from tree # 15 - **PLOT15ST** — numeric(1), range 1.0000 57.0000, number Number of stumps in a density plot centered at a random distance and direction from tree # 15 - **PLOT35TR** — numeric(1), range 3.0000 39.0000, number Number of trees in a density plot centered at a random distance and direction from tree # 35 - **PLOT35ST** — numeric(1), range 3.0000 61.0000, number Number of stumps in a density plot centered at a random distance and direction from tree # 15 - **PLOTSIZE** — numeric(0.010000), range 0.0100 0.1000, hectares Area of each circular density plot - **CALCDENS** — numeric(0.010000), range 123.5500 8895.4805, number Density as calculated by T. Bell ## Entity 2: TP08802 ### Attribute list - **DATACODE** — varchar, freetext Data set code - **FORMAT** — numeric(1), range 2.0000 2.0000, number Format number - **SITE** — varchar, enum Site identification code - **L701** — UNIT L701 - **L107** — UNIT L107 - **L111** — UNIT L111 - **L405** — UNIT L405 - **DENSITY** — varchar, enum Stocking density level - **C** — Control;no thinning within the stand - **M** — Medium;density was reduced to 25% of maximum stand density index - **L** — Low;density was reduced to 15% of maximum stand density index - **TMT** — varchar, enum Type of treatment applied to plots within stands - **F** — Fertilized - **PF** — Fertilized and pruned - **C** — Control - **P** — Pruned - **GROWTHYR** — numeric(1), range 1981.0000 81.0000, year (yyyy) Growth year sample was taken - **YEAR** — numeric(1), range 1981.0000 1996.0000, year (yyyy) Year of data collection - **MONTH** — numeric(1), range 4.0000 12.0000, month of year Month of data collection - **DAY** — numeric(1), range 1.0000 31.0000, day of month Day of data collection - **TREENO** — numeric(1), range 1.0000 290.0000, number Permanent tree tag number - **SPECIES** — varchar, enum Species code according to Garrison et al., 1972 - **ABAM** — Abies amabilis - **ABPR** — Abies procera - **PSME** — Pseudotsuga menziesii - **THPL** — Thuja plicata - **TSHE** — Tsuga heterophylla - **STATE** — varchar, enum Tree condition - **1** — Dead - **** — Alive - **DBH** — numeric(0.100000), range 0.4000 347.0000, centimeters Diameter at breast height - **HEIGHT** — numeric(0.100000), range 15.0000 997.0000, meters Total tree height - **BARK1** — numeric(1), range 1.0000 30.0000, millimeters Bark thickness direction 1 - **GR1** — numeric(1), range 1.0000 22.0000, millimeters Growth increment of last 2 years, direction 1 - **PRE1** — numeric(1), range 4.0000 55.0000, millimeters Pre-thinning ring measurements together (rings 4 - 8), direction 1 - **TOT1** — numeric(1), range 13.0000 92.0000, millimeters Thickness of last 10 years growth, direction 1 - **SAP1** — numeric(1), range 8.0000 85.0000, millimeters Sapwood thickness direction 1 - **BARK2** — numeric(1), range 2.0000 51.0000, millimeters Bark thickness direction 2 - **GR2** — numeric(1), range 1.0000 25.0000, millimeters Growth increment of last 2 years, direction 2 - **PRE2** — numeric(1), range 4.0000 55.0000, millimeters Pre-thinning ring measurements together (rings 4 - 8), direction 2 - **TOT2** — numeric(1), range 18.0000 116.0000, millimeters Thickness of last 10 years growth, direction 2 - **SAP2** — numeric(1), range 5.0000 90.0000, millimeters Sapwood thickness direction 2 - **FLAGBAD** — varchar, enum Indicates whether diameter value may be a field error - ** ** — Diameter value appears to be error-free - ***** — If present then the diameter value is suspected of being a field error - **MEASDATE** — varchar, enum Indicates whether height was measured in a different year - ** ** — Height measured on date listed in the data file - ***** — Var 'height' measured in 1982, refer to file tp8803 for date - **COND** — varchar, enum Tree condition code - **6** — Leaning - **7** — Animal damage to base (scars) - **3** — Snow bent - **5** — Small tree, broken top, probably will live - **8** — Animal damage to crown - **2** — Broken top-dead - **4** — Phellinus infection - **NOTES** — varchar, freetext Comments ## Entity 3: TP08803 ### Attribute list - **DATACODE** — varchar, freetext Data set code - **FORMAT** — numeric(1), range 3.0000 3.0000, number Format number - **SITE** — varchar, enum Site identification code - **L701** — UNIT L701 - **L107** — UNIT L107 - **L111** — UNIT L111 - **L405** — UNIT L405 - **DENSITY** — varchar, enum Stocking density level - **C** — Control;no thinning within the stand - **M** — Medium;density was reduced to 25% of maximum stand density index - **L** — Low;density was reduced to 15% of maximum stand density index - **TMT** — varchar, enum Type of treatment applied to plots within stands - **F** — Fertilized - **PF** — Fertilized and pruned - **C** — Control - **P** — Pruned - **GROWTHYR** — numeric(1), range 1981.0000 1981.0000, year (yyyy) Growth year sample was taken - **YEAR** — numeric(1), range 1981.0000 1982.0000, year (yyyy) Year of data collection - **MONTH** — numeric(1), range 5.0000 9.0000, month of year Month of data collection - **DAY** — numeric(1), range 2.0000 28.0000, day of month Day of data collection - **OLDNO** — numeric(1), range 1.0000 200.0000, number Tree tag number originally assigned - **TREENO** — numeric(1), range 1.0000 286.0000, number Permanent tree tag number - **STATUS** — varchar, enum Code indicating whether tree was alive, cut, or delayed height measurement - ** ** — Alive - **H** — Alive, height measured on date listed-dbh measured at a dif- ferent date listed in file tp8802 - **C** — Cut after initially being numbered in 1981-does not appear in data set ## Entity 4: TP08804 ### Attribute list - **DATACODE** — varchar, freetext Data set code - **FORMAT** — numeric(1), range 4.0000 4.0000, number Format number - **SITE** — varchar, enum Site identification code - **L701** — UNIT L701 - **L107** — UNIT L107 - **L111** — UNIT L111 - **L405** — UNIT L405 - **DENSITY** — varchar, enum Stocking density level - **C** — Control;no thinning within the stand - **M** — Medium;density was reduced to 25% of maximum stand density index - **L** — Low;density was reduced to 15% of maximum stand density index - **TMT** — varchar, enum Type of treatment applied to plots within stands - **F** — Fertilized - **PF** — Fertilized and pruned - **C** — Control - **P** — Pruned - **YEAR** — numeric(1), range 1983.0000 1987.0000, year (yyyy) Year of data collection - **MONTH** — numeric(1), range 10.0000 11.0000, month of year Month of data collection - **DAY** — numeric(1), range 1.0000 28.0000, day of month Day of data collection - **SPECIES** — varchar, enum Species code according to Garrison et al., 1972 - **ABAM** — Abies amabilis - **ABPR** — Abies procera - **PSME** — Pseudotsuga menziesii - **THPL** — Thuja plicata - **TSHE** — Tsuga heterophylla - **DBH** — numeric(0.100000), range 1.9000 124.0000, centimeters Diameter at breast height - **SAP1** — numeric(1), range 5.0000 36.0000, millimeters Sapwood thickness direction 1 - **GR1** — numeric(1), range 1.0000 13.0000, millimeters Growth increment of last 2 years, direction 1 - **PRE1** — numeric(1), range 5.0000 29.0000, millimeters Pre-thinning ring measurements together (rings 4 - 8), direction 1 - **SAP2** — numeric(1), range 6.0000 38.0000, millimeters Sapwood thickness direction 2 - **GR2** — numeric(1), range 2.0000 13.0000, millimeters Growth increment of last 2 years, direction 2 - **PRE2** — numeric(1), range 9.0000 36.0000, millimeters Pre-thinning ring measurements together (rings 4 - 8), direction 2 - **BARK1** — numeric(1), range 1.0000 6.0000, millimeters Bark thickness direction 1 - **BARK2** — numeric(1), range 1.0000 5.0000, millimeters Bark thickness direction 2 - **TOT1** — numeric(1), range 7.0000 65.0000, millimeters Thickness of last 10 years growth, direction 1 - **TOT2** — numeric(1), range 16.0000 80.0000, millimeters Thickness of last 10 years growth, direction 2 - **AGE_DBH5** — numeric(1), range 5.0000 10.0000, year (yyyy) Age at dbh 5 ## Entity 5: TP08805 ### Attribute list - **DATACODE** — varchar, freetext Data set code - **FORMAT** — numeric(1), range 5.0000 5.0000, number Format number - **SITE** — varchar, enum Site identification code - **L701** — UNIT L701 - **L107** — UNIT L107 - **L111** — UNIT L111 - **L405** — UNIT L405 - **DENSITY** — varchar, enum Stocking density level - **C** — Control;no thinning within the stand - **M** — Medium;density was reduced to 25% of maximum stand density index - **L** — Low;density was reduced to 15% of maximum stand density index - **TMT** — varchar, enum Type of treatment applied to plots within stands - **F** — Fertilized - **PF** — Fertilized and pruned - **C** — Control - **P** — Pruned - **COVTALL** — numeric(1), range 0.0000 90.0000, number Tall cover: number of trees competing with at least 10% of the crowns of study trees - **NFIX1** — varchar, enum Indicates whether competing trees are N-fixers - ** ** — Not nitrogen fixer - **F** — Nitrogen fixer - **COVMED** — numeric(1), range 5.0000 100.0000, number Medium cover: number of shrubs, stumps, or sprouts at shoulder height competing with the study trees - **NFIX2** — varchar, enum Indicates whether competing shrubs, stumps or sprouts are N-fixers - ** ** — Not nitrogen fixer - **F** — Nitrogen fixer - **COVLOW** — numeric(1), range 5.0000 100.0000, number Low cover: number of ground-cover plants lower than knee-height competing with the study trees - **NFIX3** — varchar, enum Indicates whether ground-cover plants are N-fixers - ** ** — Not nitrogen fixer - **F** — Nitrogen fixer ## Entity 6: TP08806 ### Attribute list - **DATACODE** — varchar, freetext Data set code - **FORMAT** — numeric(1), range 6.0000 6.0000, number Format number - **SITE** — varchar, enum Site identification code - **L701** — UNIT L701 - **L107** — UNIT L107 - **L111** — UNIT L111 - **L405** — UNIT L405 - **DENSITY** — varchar, enum Stocking density level - **C** — Control;no thinning within the stand - **M** — Medium;density was reduced to 25% of maximum stand density index - **L** — Low;density was reduced to 15% of maximum stand density index - **TMT** — varchar, enum Type of treatment applied to plots within stands - **F** — Fertilized - **PF** — Fertilized and pruned - **C** — Control - **P** — Pruned - **WETWTINT** — numeric(0.001000), range 20.0000 20.9900, grams Net weight of soil before incubation - **MOIST** — numeric(1), range 25.0000 41.0000, percent Percent moisture of soil - **NH4WET** — numeric(0.100000), range 0.2000 5.9000, parts per million Ammonia concentration in solution extracted from soil - **SOILINIT** — numeric(0.010000), range 0.0000 27.9200, grams Ammonia concentration in the soil adjusted for weight, water - **WTWTINCV** — numeric(0.001000), range 20.0000 20.9800, parts per million Wet weight of soil before incubation - **NH4INC** — numeric(0.100000), range 6.7000 25.5000, parts per million Ammonia concentration in solution extracted from soil after incubation - **SOILINC** — numeric(0.010000), range 24.8400 108.1000, parts per million Ammonia concentration in soil adjusted for weight, water after incubation - **INIBLANC** — numeric(0.100000), range 0.1000 0.4000, parts per million Ammonia concentration in blank vial solutions - **INCBLANC** — numeric(0.100000), range 0.3000 0.8000, parts per million Ammonia concentration in blank vial solutions for incubations ## Entity 7: TP08807 ### Attribute list - **DATACODE** — varchar, freetext Data set code - **FORMAT** — numeric(1), range 7.0000 7.0000, number Format number - **SITE** — varchar, enum Site identification code - **L701** — UNIT L701 - **L107** — UNIT L107 - **L111** — UNIT L111 - **L405** — UNIT L405 - **DENSITY** — varchar, enum Stocking density level - **C** — Control;no thinning within the stand - **M** — Medium;density was reduced to 25% of maximum stand density index - **L** — Low;density was reduced to 15% of maximum stand density index - **TMT** — varchar, enum Type of treatment applied to plots within stands - **F** — Fertilized - **PF** — Fertilized and pruned - **C** — Control - **P** — Pruned - **YEAR** — numeric(1), range 1996.0000 1996.0000, year (yyyy) Year of data collection - **TOTN** — numeric(0.010000), range 0.8600 1.2000, percent Total N - **TOTP** — numeric(0.010000), range 0.1600 0.2800, percent Total P - **K** — numeric(0.001000), range 0.6570 1.0740, percent K - **CA** — numeric(0.001000), range 0.3580 0.6660, percent Ca - **MG** — numeric(0.001000), range 0.1010 0.1710, percent Mg ## Attribute Definitions **AGE_DBH5** Age at dbh 5 **BARK1** Bark thickness direction 1 **BARK2** Bark thickness direction 2 **CA** Ca **CALCDENS** Density as calculated by T. Bell **COND** Tree condition code **COVLOW** Low cover: number of ground-cover plants lower than knee-height competing with the study trees **COVMED** Medium cover: number of shrubs, stumps, or sprouts at shoulder height competing with the study trees **COVTALL** Tall cover: number of trees competing with at least 10% of the crowns of study trees **DATACODE** Data set code **DAY** Day of data collection **DBH** Diameter at breast height **DENSITY** Stocking density level **FLAGBAD** Indicates whether diameter value may be a field error **FORMAT** Format number **GR1** Growth increment of last 2 years, direction 1 **GR2** Growth increment of last 2 years, direction 2 **GROWTHYR** Growth year sample was taken **HEIGHT** Total tree height **INCBLANC** Ammonia concentration in blank vial solutions for incubations **INIBLANC** Ammonia concentration in blank vial solutions **K** K **MEASDATE** Indicates whether height was measured in a different year **MG** Mg **MOIST** Percent moisture of soil **MONTH** Month of data collection **NFIX1** Indicates whether competing trees are N-fixers **NFIX2** Indicates whether competing shrubs, stumps or sprouts are N-fixers **NFIX3** Indicates whether ground-cover plants are N-fixers **NH4INC** Ammonia concentration in solution extracted from soil after incubation **NH4WET** Ammonia concentration in solution extracted from soil **NOTES** Comments **OLDNO** Tree tag number originally assigned **PLOT15ST** Number of stumps in a density plot centered at a random distance and direction from tree # 15 **PLOT15TR** Number of trees in a density plot centered at a random distance and direction from tree # 15 **PLOT35ST** Number of stumps in a density plot centered at a random distance and direction from tree # 15 **PLOT35TR** Number of trees in a density plot centered at a random distance and direction from tree # 35 **PLOTSIZE** Area of each circular density plot **PRE1** Pre-thinning ring measurements together (rings 4 - 8), direction 1 **PRE2** Pre-thinning ring measurements together (rings 4 - 8), direction 2 **SAP1** Sapwood thickness direction 1 **SAP2** Sapwood thickness direction 2 **SITE** Site identification code **SOILINC** Ammonia concentration in soil adjusted for weight, water after incubation **SOILINIT** Ammonia concentration in the soil adjusted for weight, water **SPECIES** Species code according to Garrison et al., 1972 **STATE** Tree condition **STATUS** Code indicating whether tree was alive, cut, or delayed height measurement **TMT** Type of treatment applied to plots within stands **TOT1** Thickness of last 10 years growth, direction 1 **TOT2** Thickness of last 10 years growth, direction 2 **TOTN** Total N **TOTP** Total P **TREENO** Permanent tree tag number **WETWTINT** Net weight of soil before incubation **WTWTINCV** Wet weight of soil before incubation **YEAR** Year of data collection ## Enumerated Domains ### SITE - **L701** — UNIT L701 - **L107** — UNIT L107 - **L111** — UNIT L111 - **L405** — UNIT L405 ### DENSITY - **C** — Control;no thinning within the stand - **M** — Medium;density was reduced to 25% of maximum stand density index - **L** — Low;density was reduced to 15% of maximum stand density index ### TMT - **F** — Fertilized - **PF** — Fertilized and pruned - **C** — Control - **P** — Pruned ### SPECIES - **ABAM** — Abies amabilis - **ABPR** — Abies procera - **PSME** — Pseudotsuga menziesii - **THPL** — Thuja plicata - **TSHE** — Tsuga heterophylla ### STATE - **1** — Dead - **** — Alive ### FLAGBAD - ** ** — Diameter value appears to be error-free - ***** — If present then the diameter value is suspected of being a field error ### MEASDATE - ** ** — Height measured on date listed in the data file - ***** — Var 'height' measured in 1982, refer to file tp8803 for date ### COND - **6** — Leaning - **7** — Animal damage to base (scars) - **3** — Snow bent - **5** — Small tree, broken top, probably will live - **8** — Animal damage to crown - **2** — Broken top-dead - **4** — Phellinus infection ### STATUS - ** ** — Alive - **H** — Alive, height measured on date listed-dbh measured at a dif- ferent date listed in file tp8802 - **C** — Cut after initially being numbered in 1981-does not appear in data set ### NFIX1 - ** ** — Not nitrogen fixer - **F** — Nitrogen fixer ### NFIX2 - ** ** — Not nitrogen fixer - **F** — Nitrogen fixer ### NFIX3 - ** ** — Not nitrogen fixer - **F** — Nitrogen fixer