# Species interactions during succession in the western Cascade Range of Oregon, 1990 to present

## Dataset

- **Database Code:** TP103

- **Title:** Species interactions during succession in the western Cascade Range of Oregon, 1990 to present

- **Abstract:**

The factors that contribute to plant species establishment and decline following disturbance determine the rates and patterns of successional change of a system. In this long-term field experiment, we test the commonly held assumption that competition for space or resources by dominant species determines the outcome of succession. Specifically, we examine the population- and community-level consequences of removing one or more potentially dominant species from the post-disturbance community after clearcut logging and burning of a mature/old-growth Douglas-fir forest. Experimental treatments include: (1) removal of early-seral annual, Senecio sylvaticus, or perennial, Epilobium angustifolium—or both—to test the influences of these early-seral dominants on subsequent community development; (2) removal of all species except Senecio or Epilobium, to test whether the decline of these early-seral dominants is driven by competitive displacement; or (3) removal of shade-tolerant forest species that dominate subsequent stages of succession—Rubus ursinus or Berberis nervosa plus Gaultheria shallon—to test the influences of these long-lived perennials on understory development. The experiment is a randomized block design comprising eight removal treatments plus a control replicated in each of 25 blocks. Removal (reduction in competition) is achieved by removing seedlings or vegetative stems annually from a treatment area of 2.5 x 2.5 m. Sample plots (1 x 1 m) centered within these are used to estimate cover of all vascular plant species and, for the first 8 yr of the experiment, stem density and height, facilitating estimates of above-ground biomass. Pre-harvest data were collected in 1990, logging/burning occurred in 1991, and removal treatments and post-treatment sampling were initiated in 1992. Six of the nine experimental treatments were terminated between 1996 and 1998, with loss of early-seral Senecio and Epilobium from the system. The remaining three treatments (removal of Rubus, removal of Berberis plus Gaultheria, and the control) have been sampled annually since 1992. All regeneration of trees within the experimental area has occurred naturally. In September 2018, the study site was subjected to a controlled burn to limit spread of the Terwilliger Fire from the north. In 2019, plots were assessed for burn severity. Species' removals were terminated, but cover measurements continue in the plots.


- **Keywords:** community dynamics; trophic structure; succession; primary production; species interactions; competition; disturbance; burning; timber harvest; clearcutting; Long-Term Ecological Research (LTER); primary production; disturbance

- **Temporal coverage:** 1990-07-01 to 2024-07-03

- **Principal Investigator:** Charles B. Halpern


## Data Entities

1. Species Cover, Density, and Biomass Measurement Data
2. Species Removals (counts)
3. Species Cover, Density, and Biomass Measurements Original Fieldsheet Format
4. Species Cover, Density, and Biomass Measurements Modified Fieldsheet Format
5. Equations used for calculating biomass
6. Estimated biomass using measurement data and equations
7. Assessment of 2018 burn severity

## Entity 1: Species Cover, Density, and Biomass Measurement Data

### Attribute list

- **DBCODE** — varchar, enum

Database code

  - **TP103** — FSDB database code TP103


- **ENTITY** — numeric(1.000000), range  1.0000  1.0000, number

Entity number


- **STUDY** — varchar, enum

Study site location

  - **Starrbright** — Starrbright Timber Sale


- **YEAR** — numeric(1.000000), range  1990.0000  2024.0000, number

Sampling year


- **SDATE** — datetime, freetext

Vegetation sampling date


- **SEASON** — varchar, enum

Sampling season (1 - 3)

  - **U** — Unknown season of sampling
  - **1** — First season of sampling per year (usually june)
  - **2** — Second season of sampling per year (usually july)
  - **3** — Third season of sampling per year (usually august)


- **BLOCK** — numeric(1.000000), range  1.0000  25.0000, number

Sample block (1-25)


- **PLOT** — numeric(1.000000), range  1.0000  9.0000, number

Sample plot w/in block (1-9)


- **TMT** — varchar, enum

Treatment code (A-I)

  - **A** — Control
  - **B** — Senecio sylvaticus removed
  - **C** — Epilobium angustifolium removed
  - **D** — Senecio sylvaticus and Epilobium angustifolium removed
  - **E** — Senecio sylvaticus and community removed
  - **F** — Epilobium angustifolium and community removed
  - **G** — Community removed
  - **H** — Rubus ursinus removed
  - **I** — Berberis nervosa and Gaultheria shallon removed


- **SPECIES** — varchar, enum

Species acronym according to Garrison et al. 1974

  - **BORAGI** — Boraginaceae
  - **LILIAC** — Liliaceae
  - **ABGR** — Abies grandis
  - **ACCI** — Acer circinatum
  - **ACMA** — Acer macrophyllum
  - **ACTR** — Achlys triphylla
  - **ADBI** — Adenocaulon bicolor
  - **AGGR** — Agoseris grandiflora
  - **AGHE** — Agoseris heterophylla
  - **AGROS** — Agrostis
  - **AGEX** — Agrostis exarata
  - **AGTE** — Agrostis tenuis
  - **AICA** — Aira caryophyllea
  - **AMAL** — Amelanchier alnifolia
  - **ANMA** — Anaphalis margaritacea
  - **ANDE** — Anemone deltoidea
  - **ANLY2** — Anemone lyallii
  - **ARME** — Arbutus menziesii
  - **ARCO3** — Arctostaphylos columbiana
  - **AREL** — Arrhenatherum elatius
  - **ASCA3** — Asarum caudatum
  - **BENE** — Berberis nervosa
  - **BROMU** — Bromus
  - **BRVU** — Bromus vulgaris
  - **CASC2** — Campanula scouleri
  - **CARDA** — Cardamine
  - **CAOL** — Cardamine oligosperma
  - **CAAM** — Carex amplifolia
  - **CEANO** — Ceanothus
  - **CEIN** — Ceanothus integerrimus
  - **CESA** — Ceanothus sanguineus
  - **CEVE** — Ceanothus velutinus
  - **CERAS** — Cerastium
  - **CEVU** — Cerastium vulgatum
  - **CHME** — Chimaphila menziesii
  - **CHUM** — Chimaphila umbellata
  - **CHLE2** — Chrysanthemum leucanthemum
  - **CIRSI** — Cirsium
  - **CIAR** — Cirsium arvense
  - **CIVU** — Cirsium vulgare
  - **CLUN** — Clintonia uniflora
  - **COHE** — Collomia heterophylla
  - **COCA2** — Conyza canadensis
  - **CONU** — Cornus nuttallii
  - **CRCA** — Crepis capillaris
  - **DEEL** — Deschampsia elongata
  - **ELGL** — Elymus glaucus
  - **EPILO** — Epilobium
  - **EPMI** — Epilobium minutum
  - **EPPA** — Epilobium paniculatum
  - **EPWA** — Epilobium watsonii
  - **EQTE** — Equisetum telmateia
  - **FESTU** — Festuca
  - **FEMY** — Festuca myuros
  - **FEOC** — Festuca occidentalis
  - **FRVE** — Fragaria vesca
  - **GALIU** — Galium
  - **GATR** — Galium triflorum
  - **GAFR** — Garrya fremontii
  - **GASH** — Gaultheria shallon
  - **GNPU** — Gnaphalium purpureum
  - **GNUL** — Gnaphalium uliginosum
  - **GOOB** — Goodyera oblongifolia
  - **HIAL** — Hieracium albiflorum
  - **HOLA** — Holcus lanatus
  - **HODI** — Holodiscus discolor
  - **HYPE** — Hypericum perforatum
  - **HYRA** — Hypochaeris radicata
  - **LAMU** — Lactuca muralis
  - **LASE** — Lactuca serriola
  - **LIBO2** — Linnaea borealis
  - **LISTE** — Listera
  - **LICO3** — Listera cordata
  - **LOCI** — Lonicera ciliosa
  - **LOCR** — Lotus crassifolius
  - **LOMI** — Lotus micranthus
  - **LOPU** — Lotus purshianus
  - **LULA** — Lupinus latifolius
  - **MAGR** — Madia gracilis
  - **MOPE** — Montia perfoliata
  - **MOSI** — Montia sibirica
  - **MYDI** — Myosotis discolor
  - **NEPA** — Nemophila parviflora
  - **OSCH** — Osmorhiza chilensis
  - **PEFR2** — Petasites frigidus
  - **PILA** — Pinus lambertiana
  - **POLYG** — Polygonum
  - **PODO** — Polygonum douglasii
  - **POMU** — Polystichum munitum
  - **PRVU** — Prunella vulgaris
  - **PREM** — Prunus emarginata
  - **PSME** — Pseudotsuga menziesii
  - **PTAQ** — Pteridium aquilinum
  - **PYAS** — Pyrola asarifolia
  - **PYPI** — Pyrola picta
  - **RAUN2** — Ranunculus uncinatus
  - **RHMA** — Rhododendron macrophyllum
  - **RIBES** — Ribes
  - **RILO** — Ribes lobbii
  - **ROGY** — Rosa gymnocarpa
  - **RUBUS** — Rubus
  - **RULE** — Rubus leucodermis
  - **RUNI** — Rubus nivalis
  - **RUPA** — Rubus parviflorus
  - **RUUR** — Rubus ursinus
  - **SACE** — Sambucus cerulea
  - **SEJA** — Senecio jacobaea
  - **SESY** — Senecio sylvaticus
  - **SIAN2** — Silene antirrhina
  - **SOAR** — Sonchus arvensis
  - **STELL** — Stellaria
  - **STCA2** — Stellaria calycantha
  - **TABR** — Taxus brevifolia
  - **THPL** — Thuja plicata
  - **TITR** — Tiarella trifoliata
  - **TRLA2** — Trientalis latifolia
  - **TROV** — Trillium ovatum
  - **TRCA** — Trisetum canescens
  - **TRCE** — Trisetum cernuum
  - **TSHE** — Tsuga heterophylla
  - **VAME** — Vaccinium membranaceum
  - **VAPA** — Vaccinium parvifolium
  - **VAHE** — Vancouveria hexandra
  - **VEAR** — Veronica arvensis
  - **VESE** — Veronica serpyllifolia
  - **VISE** — Viola sempervirens
  - **WHMO** — Whipplea modesta
  - **CACH** — Castanopsis chrysophylla
  - **EBAU** — Eburophyton austiniae
  - **SADO** — Satureja douglasii
  - **SMST** — Smilacina stellata
  - **EPAN** — Epilobium angustifolium
  - **SYMO** — Symphoricarpos mollis
  - **GNMI** — Gnaphalium microcephalum
  - **LUCA2** — Luzula campestris
  - **UNKN** — unknown plant
  - **HERB** — Herb
  - **TREE** — Tree
  - **SHRUB** — Shrub
  - **MOSS** — Moss
  - **LICHEN** — Lichen
  - **GRASS** — Grass
  - **LITTER** — Litter
  - **CWD** — Coarse woody debris
  - **BARE** — Bare ground
  - **LOG** — Log
  - **STONE** — Stone
  - **BUTT** — Tree butt
  - **STUMP** — Tree stump
  - **SNAG** — Snag
  - **UNKN2** — unknown plant 2
  - **ROOT** — Root


- **STRATUM** — varchar, enum

Pre-harvest tree stratum: (O)verstory, (S)ubcanopy, (U)nderstory

  - **O** — Overstory layer
  - **S** — Subcanopy layer
  - **U** — Understory layer
  - **N** — Not applicable; STRATUM applies only to tree species in the pre-harvest sample (1990)


- **COV1_4TH** — numeric(0.100000), range  0.0000  100.0000, percent

Cover in the "quarter" plot (0.25 x 0.25 m), used to subsample when species density was high


- **COVER** — numeric(0.100000), range  0.0000  100.0000, percent

Cover in the 1 m2 plot


- **DENSITYS** — numeric(1.000000), range  0.0000  3300.0000, number

Density of seedlings (1 m2 plot)


- **DENSITYM** — numeric(1.000000), range  0.0000  105.0000, number

Density of mature stems/sprouts (1 m2 plot)


- **DBA** — numeric(0.100000), range  0.0000  100.0000, centimeters

Basal diameter


- **HT** — numeric(0.100000), range  0.0000  145.0000, centimeters

Total height (including inflorescence if present)


- **HTV** — numeric(0.100000), range  0.0000  330.0000, centimeters

Height of vegetative portion


- **REC_NO** — numeric(1.000000), range  1.0000  1135.0000, number

Unique record number for a given year, season, block, plot, tmt, species and stratum


## Entity 2: Species Removals (counts)

### Attribute list

- **DBCODE** — varchar, enum

Database code

  - **TP103** — FSDB database code TP103


- **ENTITY** — numeric(1.000000), range  2.0000  2.0000, number

Entity number


- **STUDY** — varchar, enum

Study site location

  - **Starrbright** — Starrbright Timber Sale


- **SDATE** — datetime, freetext

Vegetation sampling date


- **RDATE** — datetime, freetext

Date of species removal (yyyymmdd)


- **SEASON** — varchar, enum

Sampling season (1 - 3)

  - **U** — Unknown season of sampling
  - **1** — First season of sampling per year (usually june)
  - **2** — Second season of sampling per year (usually july)
  - **3** — Third season of sampling per year (usually august)


- **BLOCK** — numeric(1.000000), range  1.0000  25.0000, number

Sample block (1-25)


- **PLOT** — numeric(1.000000), range  1.0000  9.0000, number

Sample plot w/in block (1-9)


- **TMT** — varchar, enum

Treatment code (A-I)

  - **A** — Control
  - **B** — Senecio sylvaticus removed
  - **C** — Epilobium angustifolium removed
  - **D** — Senecio sylvaticus and Epilobium angustifolium removed
  - **E** — Senecio sylvaticus and community removed
  - **F** — Epilobium angustifolium and community removed
  - **G** — Community removed
  - **H** — Rubus ursinus removed
  - **I** — Berberis nervosa and Gaultheria shallon removed


- **SPECIES** — varchar, enum

Species acronym according to Garrison et al. 1974

  - **BORAGI** — Boraginaceae
  - **LILIAC** — Liliaceae
  - **ABGR** — Abies grandis
  - **ACCI** — Acer circinatum
  - **ACMA** — Acer macrophyllum
  - **ACTR** — Achlys triphylla
  - **ADBI** — Adenocaulon bicolor
  - **AGGR** — Agoseris grandiflora
  - **AGHE** — Agoseris heterophylla
  - **AGROS** — Agrostis
  - **AGEX** — Agrostis exarata
  - **AGTE** — Agrostis tenuis
  - **AICA** — Aira caryophyllea
  - **AMAL** — Amelanchier alnifolia
  - **ANMA** — Anaphalis margaritacea
  - **ANDE** — Anemone deltoidea
  - **ANLY2** — Anemone lyallii
  - **ARME** — Arbutus menziesii
  - **ARCO3** — Arctostaphylos columbiana
  - **AREL** — Arrhenatherum elatius
  - **ASCA3** — Asarum caudatum
  - **BENE** — Berberis nervosa
  - **BROMU** — Bromus
  - **BRVU** — Bromus vulgaris
  - **CASC2** — Campanula scouleri
  - **CARDA** — Cardamine
  - **CAOL** — Cardamine oligosperma
  - **CAAM** — Carex amplifolia
  - **CEANO** — Ceanothus
  - **CEIN** — Ceanothus integerrimus
  - **CESA** — Ceanothus sanguineus
  - **CEVE** — Ceanothus velutinus
  - **CERAS** — Cerastium
  - **CEVU** — Cerastium vulgatum
  - **CHME** — Chimaphila menziesii
  - **CHUM** — Chimaphila umbellata
  - **CHLE2** — Chrysanthemum leucanthemum
  - **CIRSI** — Cirsium
  - **CIAR** — Cirsium arvense
  - **CIVU** — Cirsium vulgare
  - **CLUN** — Clintonia uniflora
  - **COHE** — Collomia heterophylla
  - **COCA2** — Conyza canadensis
  - **CONU** — Cornus nuttallii
  - **CRCA** — Crepis capillaris
  - **DEEL** — Deschampsia elongata
  - **ELGL** — Elymus glaucus
  - **EPILO** — Epilobium
  - **EPMI** — Epilobium minutum
  - **EPPA** — Epilobium paniculatum
  - **EPWA** — Epilobium watsonii
  - **EQTE** — Equisetum telmateia
  - **FESTU** — Festuca
  - **FEMY** — Festuca myuros
  - **FEOC** — Festuca occidentalis
  - **FRVE** — Fragaria vesca
  - **GALIU** — Galium
  - **GATR** — Galium triflorum
  - **GAFR** — Garrya fremontii
  - **GASH** — Gaultheria shallon
  - **GNPU** — Gnaphalium purpureum
  - **GNUL** — Gnaphalium uliginosum
  - **GOOB** — Goodyera oblongifolia
  - **HIAL** — Hieracium albiflorum
  - **HOLA** — Holcus lanatus
  - **HODI** — Holodiscus discolor
  - **HYPE** — Hypericum perforatum
  - **HYRA** — Hypochaeris radicata
  - **LAMU** — Lactuca muralis
  - **LASE** — Lactuca serriola
  - **LIBO2** — Linnaea borealis
  - **LISTE** — Listera
  - **LICO3** — Listera cordata
  - **LOCI** — Lonicera ciliosa
  - **LOCR** — Lotus crassifolius
  - **LOMI** — Lotus micranthus
  - **LOPU** — Lotus purshianus
  - **LULA** — Lupinus latifolius
  - **MAGR** — Madia gracilis
  - **MOPE** — Montia perfoliata
  - **MOSI** — Montia sibirica
  - **MYDI** — Myosotis discolor
  - **NEPA** — Nemophila parviflora
  - **OSCH** — Osmorhiza chilensis
  - **PEFR2** — Petasites frigidus
  - **PILA** — Pinus lambertiana
  - **POLYG** — Polygonum
  - **PODO** — Polygonum douglasii
  - **POMU** — Polystichum munitum
  - **PRVU** — Prunella vulgaris
  - **PREM** — Prunus emarginata
  - **PSME** — Pseudotsuga menziesii
  - **PTAQ** — Pteridium aquilinum
  - **PYAS** — Pyrola asarifolia
  - **PYPI** — Pyrola picta
  - **RAUN2** — Ranunculus uncinatus
  - **RHMA** — Rhododendron macrophyllum
  - **RIBES** — Ribes
  - **RILO** — Ribes lobbii
  - **ROGY** — Rosa gymnocarpa
  - **RUBUS** — Rubus
  - **RULE** — Rubus leucodermis
  - **RUNI** — Rubus nivalis
  - **RUPA** — Rubus parviflorus
  - **RUUR** — Rubus ursinus
  - **SACE** — Sambucus cerulea
  - **SEJA** — Senecio jacobaea
  - **SESY** — Senecio sylvaticus
  - **SIAN2** — Silene antirrhina
  - **SOAR** — Sonchus arvensis
  - **STELL** — Stellaria
  - **STCA2** — Stellaria calycantha
  - **TABR** — Taxus brevifolia
  - **THPL** — Thuja plicata
  - **TITR** — Tiarella trifoliata
  - **TRLA2** — Trientalis latifolia
  - **TROV** — Trillium ovatum
  - **TRCA** — Trisetum canescens
  - **TRCE** — Trisetum cernuum
  - **TSHE** — Tsuga heterophylla
  - **VAME** — Vaccinium membranaceum
  - **VAPA** — Vaccinium parvifolium
  - **VAHE** — Vancouveria hexandra
  - **VEAR** — Veronica arvensis
  - **VESE** — Veronica serpyllifolia
  - **VISE** — Viola sempervirens
  - **WHMO** — Whipplea modesta
  - **CACH** — Castanopsis chrysophylla
  - **EBAU** — Eburophyton austiniae
  - **SADO** — Satureja douglasii
  - **SMST** — Smilacina stellata
  - **EPAN** — Epilobium angustifolium
  - **SYMO** — Symphoricarpos mollis
  - **GNMI** — Gnaphalium microcephalum
  - **LUCA2** — Luzula campestris
  - **UNKN** — unknown plant
  - **HERB** — Herb
  - **TREE** — Tree
  - **SHRUB** — Shrub
  - **MOSS** — Moss
  - **LICHEN** — Lichen
  - **GRASS** — Grass
  - **LITTER** — Litter
  - **CWD** — Coarse woody debris
  - **BARE** — Bare ground
  - **LOG** — Log
  - **STONE** — Stone
  - **BUTT** — Tree butt
  - **STUMP** — Tree stump
  - **SNAG** — Snag
  - **UNKN2** — unknown plant 2
  - **ROOT** — Root


- **NREMBUF** — numeric(1.000000), range  0.0000  1354.0000, number

Number of stems removed from 75 cm buffer


- **NREMPLT** — numeric(1.000000), range  0.0000  486.0000, number

Number of stems removed from 1 m2 plot


- **NOTES** — varchar, freetext

Comments


## Entity 3: Species Cover, Density, and Biomass Measurements Original Fieldsheet Format

### Attribute list

- **DBCODE** — varchar, enum

Database code

  - **TP103** — FSDB database code TP103


- **ENTITY** — numeric(1.000000), range  3.0000  3.0000, number

Entity number


- **STUDY** — varchar, enum

Study site location

  - **Starrbright** — Starrbright Timber Sale


- **YEAR** — numeric(1.000000), range  1990.0000  1996.0000, number

Sampling year


- **SDATE** — datetime, freetext

Vegetation sampling date


- **SEASON** — varchar, enum

Sampling season (1 - 3)

  - **U** — Unknown season of sampling
  - **1** — First season of sampling per year (usually june)
  - **2** — Second season of sampling per year (usually july)
  - **3** — Third season of sampling per year (usually august)


- **BLOCK** — numeric(1.000000), range  1.0000  25.0000, number

Sample block (1-25)


- **PLOT** — numeric(1.000000), range  1.0000  9.0000, number

Sample plot w/in block (1-9)


- **TMT** — varchar, enum

Treatment code (A-I)

  - **A** — Control
  - **B** — Senecio sylvaticus removed
  - **C** — Epilobium angustifolium removed
  - **D** — Senecio sylvaticus and Epilobium angustifolium removed
  - **E** — Senecio sylvaticus and community removed
  - **F** — Epilobium angustifolium and community removed
  - **G** — Community removed
  - **H** — Rubus ursinus removed
  - **I** — Berberis nervosa and Gaultheria shallon removed


- **SPECIES** — varchar, enum

Species acronym according to Garrison et al. 1974

  - **BORAGI** — Boraginaceae
  - **LILIAC** — Liliaceae
  - **ABGR** — Abies grandis
  - **ACCI** — Acer circinatum
  - **ACMA** — Acer macrophyllum
  - **ACTR** — Achlys triphylla
  - **ADBI** — Adenocaulon bicolor
  - **AGGR** — Agoseris grandiflora
  - **AGHE** — Agoseris heterophylla
  - **AGROS** — Agrostis
  - **AGEX** — Agrostis exarata
  - **AGTE** — Agrostis tenuis
  - **AICA** — Aira caryophyllea
  - **AMAL** — Amelanchier alnifolia
  - **ANMA** — Anaphalis margaritacea
  - **ANDE** — Anemone deltoidea
  - **ANLY2** — Anemone lyallii
  - **ARME** — Arbutus menziesii
  - **ARCO3** — Arctostaphylos columbiana
  - **AREL** — Arrhenatherum elatius
  - **ASCA3** — Asarum caudatum
  - **BENE** — Berberis nervosa
  - **BROMU** — Bromus
  - **BRVU** — Bromus vulgaris
  - **CASC2** — Campanula scouleri
  - **CARDA** — Cardamine
  - **CAOL** — Cardamine oligosperma
  - **CAAM** — Carex amplifolia
  - **CEANO** — Ceanothus
  - **CEIN** — Ceanothus integerrimus
  - **CESA** — Ceanothus sanguineus
  - **CEVE** — Ceanothus velutinus
  - **CERAS** — Cerastium
  - **CEVU** — Cerastium vulgatum
  - **CHME** — Chimaphila menziesii
  - **CHUM** — Chimaphila umbellata
  - **CHLE2** — Chrysanthemum leucanthemum
  - **CIRSI** — Cirsium
  - **CIAR** — Cirsium arvense
  - **CIVU** — Cirsium vulgare
  - **CLUN** — Clintonia uniflora
  - **COHE** — Collomia heterophylla
  - **COCA2** — Conyza canadensis
  - **CONU** — Cornus nuttallii
  - **CRCA** — Crepis capillaris
  - **DEEL** — Deschampsia elongata
  - **ELGL** — Elymus glaucus
  - **EPILO** — Epilobium
  - **EPMI** — Epilobium minutum
  - **EPPA** — Epilobium paniculatum
  - **EPWA** — Epilobium watsonii
  - **EQTE** — Equisetum telmateia
  - **FESTU** — Festuca
  - **FEMY** — Festuca myuros
  - **FEOC** — Festuca occidentalis
  - **FRVE** — Fragaria vesca
  - **GALIU** — Galium
  - **GATR** — Galium triflorum
  - **GAFR** — Garrya fremontii
  - **GASH** — Gaultheria shallon
  - **GNPU** — Gnaphalium purpureum
  - **GNUL** — Gnaphalium uliginosum
  - **GOOB** — Goodyera oblongifolia
  - **HIAL** — Hieracium albiflorum
  - **HOLA** — Holcus lanatus
  - **HODI** — Holodiscus discolor
  - **HYPE** — Hypericum perforatum
  - **HYRA** — Hypochaeris radicata
  - **LAMU** — Lactuca muralis
  - **LASE** — Lactuca serriola
  - **LIBO2** — Linnaea borealis
  - **LISTE** — Listera
  - **LICO3** — Listera cordata
  - **LOCI** — Lonicera ciliosa
  - **LOCR** — Lotus crassifolius
  - **LOMI** — Lotus micranthus
  - **LOPU** — Lotus purshianus
  - **LULA** — Lupinus latifolius
  - **MAGR** — Madia gracilis
  - **MOPE** — Montia perfoliata
  - **MOSI** — Montia sibirica
  - **MYDI** — Myosotis discolor
  - **NEPA** — Nemophila parviflora
  - **OSCH** — Osmorhiza chilensis
  - **PEFR2** — Petasites frigidus
  - **PILA** — Pinus lambertiana
  - **POLYG** — Polygonum
  - **PODO** — Polygonum douglasii
  - **POMU** — Polystichum munitum
  - **PRVU** — Prunella vulgaris
  - **PREM** — Prunus emarginata
  - **PSME** — Pseudotsuga menziesii
  - **PTAQ** — Pteridium aquilinum
  - **PYAS** — Pyrola asarifolia
  - **PYPI** — Pyrola picta
  - **RAUN2** — Ranunculus uncinatus
  - **RHMA** — Rhododendron macrophyllum
  - **RIBES** — Ribes
  - **RILO** — Ribes lobbii
  - **ROGY** — Rosa gymnocarpa
  - **RUBUS** — Rubus
  - **RULE** — Rubus leucodermis
  - **RUNI** — Rubus nivalis
  - **RUPA** — Rubus parviflorus
  - **RUUR** — Rubus ursinus
  - **SACE** — Sambucus cerulea
  - **SEJA** — Senecio jacobaea
  - **SESY** — Senecio sylvaticus
  - **SIAN2** — Silene antirrhina
  - **SOAR** — Sonchus arvensis
  - **STELL** — Stellaria
  - **STCA2** — Stellaria calycantha
  - **TABR** — Taxus brevifolia
  - **THPL** — Thuja plicata
  - **TITR** — Tiarella trifoliata
  - **TRLA2** — Trientalis latifolia
  - **TROV** — Trillium ovatum
  - **TRCA** — Trisetum canescens
  - **TRCE** — Trisetum cernuum
  - **TSHE** — Tsuga heterophylla
  - **VAME** — Vaccinium membranaceum
  - **VAPA** — Vaccinium parvifolium
  - **VAHE** — Vancouveria hexandra
  - **VEAR** — Veronica arvensis
  - **VESE** — Veronica serpyllifolia
  - **VISE** — Viola sempervirens
  - **WHMO** — Whipplea modesta
  - **CACH** — Castanopsis chrysophylla
  - **EBAU** — Eburophyton austiniae
  - **SADO** — Satureja douglasii
  - **SMST** — Smilacina stellata
  - **EPAN** — Epilobium angustifolium
  - **SYMO** — Symphoricarpos mollis
  - **GNMI** — Gnaphalium microcephalum
  - **LUCA2** — Luzula campestris
  - **UNKN** — unknown plant
  - **HERB** — Herb
  - **TREE** — Tree
  - **SHRUB** — Shrub
  - **MOSS** — Moss
  - **LICHEN** — Lichen
  - **GRASS** — Grass
  - **LITTER** — Litter
  - **CWD** — Coarse woody debris
  - **BARE** — Bare ground
  - **LOG** — Log
  - **STONE** — Stone
  - **BUTT** — Tree butt
  - **STUMP** — Tree stump
  - **SNAG** — Snag
  - **UNKN2** — unknown plant 2
  - **ROOT** — Root


- **STRATUM** — varchar, enum

Pre-harvest tree stratum: (O)verstory, (S)ubcanopy, (U)nderstory

  - **O** — Overstory layer
  - **S** — Subcanopy layer
  - **U** — Understory layer
  - **N** — Not applicable; STRATUM applies only to tree species in the pre-harvest sample (1990)


- **LC** — numeric(1.000000), range  1.0000  165.0000, number

Line count


- **COV1_4TH** — numeric(0.100000), range  0.0000  100.0000, percent

Cover in the "quarter" plot (0.25 x 0.25 m), used to subsample when species density was high


- **COVER** — numeric(0.100000), range  0.0000  100.0000, percent

Cover in the 1 m2 plot


- **DENSITYS** — numeric(1.000000), range  0.0000  3300.0000, number

Density of seedlings (1 m2 plot)


- **DENSITYM** — numeric(1.000000), range  0.0000  120.0000, number

Density of mature stems/sprouts (1 m2 plot)


- **STEM1BV1** — numeric(0.100000), range  0.0000  130.0000, number

Biomass parameter 1 of stem1


- **STEM1BV2** — numeric(0.100000), range  0.0000  130.0000, number

Biomass parameter 2 of stem1


- **STEM2BV1** — numeric(0.100000), range  0.0000  330.0000, number

Biomass parameter 1 of stem2


- **STEM2BV2** — numeric(0.100000), range  0.0000  150.0000, number

Biomass parameter 2 of stem2


- **STEM3BV1** — numeric(0.100000), range  0.0000  130.0000, number

Biomass parameter 1 of stem3


- **STEM3BV2** — numeric(0.100000), range  0.0000  130.0000, number

Biomass parameter 2 of stem3


- **STEM4BV1** — numeric(0.100000), range  0.0000  130.0000, number

Biomass parameter 1 of stem4


- **STEM4BV2** — numeric(0.100000), range  0.0000  130.0000, number

Biomass parameter 2 of stem4


- **STEM5BV1** — numeric(0.100000), range  0.0000  130.0000, number

Biomass parameter 1 of stem5


- **STEM5BV2** — numeric(0.100000), range  0.0000  130.0000, number

Biomass parameter 2 of stem5


## Entity 4: Species Cover, Density, and Biomass Measurements Modified Fieldsheet Format

### Attribute list

- **DBCODE** — varchar, enum

Database code

  - **TP103** — FSDB database code TP103


- **ENTITY** — numeric(1.000000), range  4.0000  4.0000, number

Entity number


- **STUDY** — varchar, enum

Study site location

  - **Starrbright** — Starrbright Timber Sale


- **YEAR** — numeric(1.000000), range  1997.0000  2002.0000, number

Sampling year


- **SDATE** — datetime, freetext

Vegetation sampling date


- **SEASON** — varchar, enum

Sampling season (1 - 3)

  - **U** — Unknown season of sampling
  - **1** — First season of sampling per year (usually june)
  - **2** — Second season of sampling per year (usually july)
  - **3** — Third season of sampling per year (usually august)


- **BLOCK** — numeric(1.000000), range  1.0000  25.0000, number

Sample block (1-25)


- **PLOT** — numeric(1.000000), range  1.0000  9.0000, number

Sample plot w/in block (1-9)


- **TMT** — varchar, enum

Treatment code (A-I)

  - **A** — Control
  - **B** — Senecio sylvaticus removed
  - **C** — Epilobium angustifolium removed
  - **D** — Senecio sylvaticus and Epilobium angustifolium removed
  - **E** — Senecio sylvaticus and community removed
  - **F** — Epilobium angustifolium and community removed
  - **G** — Community removed
  - **H** — Rubus ursinus removed
  - **I** — Berberis nervosa and Gaultheria shallon removed


- **SPECIES** — varchar, enum

Species acronym according to Garrison et al. 1974

  - **BORAGI** — Boraginaceae
  - **LILIAC** — Liliaceae
  - **ABGR** — Abies grandis
  - **ACCI** — Acer circinatum
  - **ACMA** — Acer macrophyllum
  - **ACTR** — Achlys triphylla
  - **ADBI** — Adenocaulon bicolor
  - **AGGR** — Agoseris grandiflora
  - **AGHE** — Agoseris heterophylla
  - **AGROS** — Agrostis
  - **AGEX** — Agrostis exarata
  - **AGTE** — Agrostis tenuis
  - **AICA** — Aira caryophyllea
  - **AMAL** — Amelanchier alnifolia
  - **ANMA** — Anaphalis margaritacea
  - **ANDE** — Anemone deltoidea
  - **ANLY2** — Anemone lyallii
  - **ARME** — Arbutus menziesii
  - **ARCO3** — Arctostaphylos columbiana
  - **AREL** — Arrhenatherum elatius
  - **ASCA3** — Asarum caudatum
  - **BENE** — Berberis nervosa
  - **BROMU** — Bromus
  - **BRVU** — Bromus vulgaris
  - **CASC2** — Campanula scouleri
  - **CARDA** — Cardamine
  - **CAOL** — Cardamine oligosperma
  - **CAAM** — Carex amplifolia
  - **CEANO** — Ceanothus
  - **CEIN** — Ceanothus integerrimus
  - **CESA** — Ceanothus sanguineus
  - **CEVE** — Ceanothus velutinus
  - **CERAS** — Cerastium
  - **CEVU** — Cerastium vulgatum
  - **CHME** — Chimaphila menziesii
  - **CHUM** — Chimaphila umbellata
  - **CHLE2** — Chrysanthemum leucanthemum
  - **CIRSI** — Cirsium
  - **CIAR** — Cirsium arvense
  - **CIVU** — Cirsium vulgare
  - **CLUN** — Clintonia uniflora
  - **COHE** — Collomia heterophylla
  - **COCA2** — Conyza canadensis
  - **CONU** — Cornus nuttallii
  - **CRCA** — Crepis capillaris
  - **DEEL** — Deschampsia elongata
  - **ELGL** — Elymus glaucus
  - **EPILO** — Epilobium
  - **EPMI** — Epilobium minutum
  - **EPPA** — Epilobium paniculatum
  - **EPWA** — Epilobium watsonii
  - **EQTE** — Equisetum telmateia
  - **FESTU** — Festuca
  - **FEMY** — Festuca myuros
  - **FEOC** — Festuca occidentalis
  - **FRVE** — Fragaria vesca
  - **GALIU** — Galium
  - **GATR** — Galium triflorum
  - **GAFR** — Garrya fremontii
  - **GASH** — Gaultheria shallon
  - **GNPU** — Gnaphalium purpureum
  - **GNUL** — Gnaphalium uliginosum
  - **GOOB** — Goodyera oblongifolia
  - **HIAL** — Hieracium albiflorum
  - **HOLA** — Holcus lanatus
  - **HODI** — Holodiscus discolor
  - **HYPE** — Hypericum perforatum
  - **HYRA** — Hypochaeris radicata
  - **LAMU** — Lactuca muralis
  - **LASE** — Lactuca serriola
  - **LIBO2** — Linnaea borealis
  - **LISTE** — Listera
  - **LICO3** — Listera cordata
  - **LOCI** — Lonicera ciliosa
  - **LOCR** — Lotus crassifolius
  - **LOMI** — Lotus micranthus
  - **LOPU** — Lotus purshianus
  - **LULA** — Lupinus latifolius
  - **MAGR** — Madia gracilis
  - **MOPE** — Montia perfoliata
  - **MOSI** — Montia sibirica
  - **MYDI** — Myosotis discolor
  - **NEPA** — Nemophila parviflora
  - **OSCH** — Osmorhiza chilensis
  - **PEFR2** — Petasites frigidus
  - **PILA** — Pinus lambertiana
  - **POLYG** — Polygonum
  - **PODO** — Polygonum douglasii
  - **POMU** — Polystichum munitum
  - **PRVU** — Prunella vulgaris
  - **PREM** — Prunus emarginata
  - **PSME** — Pseudotsuga menziesii
  - **PTAQ** — Pteridium aquilinum
  - **PYAS** — Pyrola asarifolia
  - **PYPI** — Pyrola picta
  - **RAUN2** — Ranunculus uncinatus
  - **RHMA** — Rhododendron macrophyllum
  - **RIBES** — Ribes
  - **RILO** — Ribes lobbii
  - **ROGY** — Rosa gymnocarpa
  - **RUBUS** — Rubus
  - **RULE** — Rubus leucodermis
  - **RUNI** — Rubus nivalis
  - **RUPA** — Rubus parviflorus
  - **RUUR** — Rubus ursinus
  - **SACE** — Sambucus cerulea
  - **SEJA** — Senecio jacobaea
  - **SESY** — Senecio sylvaticus
  - **SIAN2** — Silene antirrhina
  - **SOAR** — Sonchus arvensis
  - **STELL** — Stellaria
  - **STCA2** — Stellaria calycantha
  - **TABR** — Taxus brevifolia
  - **THPL** — Thuja plicata
  - **TITR** — Tiarella trifoliata
  - **TRLA2** — Trientalis latifolia
  - **TROV** — Trillium ovatum
  - **TRCA** — Trisetum canescens
  - **TRCE** — Trisetum cernuum
  - **TSHE** — Tsuga heterophylla
  - **VAME** — Vaccinium membranaceum
  - **VAPA** — Vaccinium parvifolium
  - **VAHE** — Vancouveria hexandra
  - **VEAR** — Veronica arvensis
  - **VESE** — Veronica serpyllifolia
  - **VISE** — Viola sempervirens
  - **WHMO** — Whipplea modesta
  - **CACH** — Castanopsis chrysophylla
  - **EBAU** — Eburophyton austiniae
  - **SADO** — Satureja douglasii
  - **SMST** — Smilacina stellata
  - **EPAN** — Epilobium angustifolium
  - **SYMO** — Symphoricarpos mollis
  - **GNMI** — Gnaphalium microcephalum
  - **LUCA2** — Luzula campestris
  - **UNKN** — unknown plant
  - **HERB** — Herb
  - **TREE** — Tree
  - **SHRUB** — Shrub
  - **MOSS** — Moss
  - **LICHEN** — Lichen
  - **GRASS** — Grass
  - **LITTER** — Litter
  - **CWD** — Coarse woody debris
  - **BARE** — Bare ground
  - **LOG** — Log
  - **STONE** — Stone
  - **BUTT** — Tree butt
  - **STUMP** — Tree stump
  - **SNAG** — Snag
  - **UNKN2** — unknown plant 2
  - **ROOT** — Root


- **LC** — numeric(1.000000), range  1.0000  10.0000, number

Line count


- **COV1_4TH** — numeric(0.100000), range  0.0000  100.0000, percent

Cover in the "quarter" plot (0.25 x 0.25 m), used to subsample when species density was high


- **COVER** — numeric(0.100000), range  0.0000  100.0000, percent

Cover in the 1 m2 plot


- **DENSITYS** — numeric(1.000000), range  0.0000  2000.0000, number

Density of seedlings (1 m2 plot)


- **DENSITYM** — numeric(1.000000), range  0.0000  70.0000, number

Density of mature stems/sprouts (1 m2 plot)


- **STEM1BV1** — numeric(0.100000), range  0.1000  150.0000, number

Biomass parameter 1 of stem1


- **STEM1BV2** — numeric(0.100000), range  0.1000  150.0000, number

Biomass parameter 2 of stem1


- **STEM1NO** — numeric(1.000000), range  1.0000  1000.0000, number

Number of stems containing the corresponding bv1 and bv2 values


- **STEM2BV1** — numeric(0.100000), range  0.1000  150.0000, number

Biomass parameter 1 of stem2


- **STEM2BV2** — numeric(0.100000), range  0.1000  150.0000, number

Biomass parameter 2 of stem2


- **STEM2NO** — numeric(1.000000), range  1.0000  1000.0000, number

Number of stems containing the corresponding bv1 and bv2 values


- **STEM3BV1** — numeric(0.100000), range  0.1000  150.0000, number

Biomass parameter 1 of stem3


- **STEM3BV2** — numeric(0.100000), range  0.1000  150.0000, number

Biomass parameter 2 of stem3


- **STEM3NO** — numeric(1.000000), range  1.0000  1000.0000, number

Number of stems containing the corresponding bv1 and bv2 values


## Entity 5: Equations used for calculating biomass

### Attribute list

- **DBCODE** — varchar, enum

Database code

  - **TP103** — FSDB database code TP103


- **ENTITY** — numeric(1.000000), range  5.0000  5.0000, number

Entity number


- **STUDY** — varchar, enum

Study site location

  - **Starrbright** — Starrbright Timber Sale


- **YEAR** — numeric(1.000000), range  1992.0000  2024.0000, number

Sampling year


- **SPECIES** — varchar, enum

Species acronym according to Garrison et al. 1974

  - **BORAGI** — Boraginaceae
  - **LILIAC** — Liliaceae
  - **ABGR** — Abies grandis
  - **ACCI** — Acer circinatum
  - **ACMA** — Acer macrophyllum
  - **ACTR** — Achlys triphylla
  - **ADBI** — Adenocaulon bicolor
  - **AGGR** — Agoseris grandiflora
  - **AGHE** — Agoseris heterophylla
  - **AGROS** — Agrostis
  - **AGEX** — Agrostis exarata
  - **AGTE** — Agrostis tenuis
  - **AICA** — Aira caryophyllea
  - **AMAL** — Amelanchier alnifolia
  - **ANMA** — Anaphalis margaritacea
  - **ANDE** — Anemone deltoidea
  - **ANLY2** — Anemone lyallii
  - **ARME** — Arbutus menziesii
  - **ARCO3** — Arctostaphylos columbiana
  - **AREL** — Arrhenatherum elatius
  - **ASCA3** — Asarum caudatum
  - **BENE** — Berberis nervosa
  - **BROMU** — Bromus
  - **BRVU** — Bromus vulgaris
  - **CASC2** — Campanula scouleri
  - **CARDA** — Cardamine
  - **CAOL** — Cardamine oligosperma
  - **CAAM** — Carex amplifolia
  - **CEANO** — Ceanothus
  - **CEIN** — Ceanothus integerrimus
  - **CESA** — Ceanothus sanguineus
  - **CEVE** — Ceanothus velutinus
  - **CERAS** — Cerastium
  - **CEVU** — Cerastium vulgatum
  - **CHME** — Chimaphila menziesii
  - **CHUM** — Chimaphila umbellata
  - **CHLE2** — Chrysanthemum leucanthemum
  - **CIRSI** — Cirsium
  - **CIAR** — Cirsium arvense
  - **CIVU** — Cirsium vulgare
  - **CLUN** — Clintonia uniflora
  - **COHE** — Collomia heterophylla
  - **COCA2** — Conyza canadensis
  - **CONU** — Cornus nuttallii
  - **CRCA** — Crepis capillaris
  - **DEEL** — Deschampsia elongata
  - **ELGL** — Elymus glaucus
  - **EPILO** — Epilobium
  - **EPMI** — Epilobium minutum
  - **EPPA** — Epilobium paniculatum
  - **EPWA** — Epilobium watsonii
  - **EQTE** — Equisetum telmateia
  - **FESTU** — Festuca
  - **FEMY** — Festuca myuros
  - **FEOC** — Festuca occidentalis
  - **FRVE** — Fragaria vesca
  - **GALIU** — Galium
  - **GATR** — Galium triflorum
  - **GAFR** — Garrya fremontii
  - **GASH** — Gaultheria shallon
  - **GNPU** — Gnaphalium purpureum
  - **GNUL** — Gnaphalium uliginosum
  - **GOOB** — Goodyera oblongifolia
  - **HIAL** — Hieracium albiflorum
  - **HOLA** — Holcus lanatus
  - **HODI** — Holodiscus discolor
  - **HYPE** — Hypericum perforatum
  - **HYRA** — Hypochaeris radicata
  - **LAMU** — Lactuca muralis
  - **LASE** — Lactuca serriola
  - **LIBO2** — Linnaea borealis
  - **LISTE** — Listera
  - **LICO3** — Listera cordata
  - **LOCI** — Lonicera ciliosa
  - **LOCR** — Lotus crassifolius
  - **LOMI** — Lotus micranthus
  - **LOPU** — Lotus purshianus
  - **LULA** — Lupinus latifolius
  - **MAGR** — Madia gracilis
  - **MOPE** — Montia perfoliata
  - **MOSI** — Montia sibirica
  - **MYDI** — Myosotis discolor
  - **NEPA** — Nemophila parviflora
  - **OSCH** — Osmorhiza chilensis
  - **PEFR2** — Petasites frigidus
  - **PILA** — Pinus lambertiana
  - **POLYG** — Polygonum
  - **PODO** — Polygonum douglasii
  - **POMU** — Polystichum munitum
  - **PRVU** — Prunella vulgaris
  - **PREM** — Prunus emarginata
  - **PSME** — Pseudotsuga menziesii
  - **PTAQ** — Pteridium aquilinum
  - **PYAS** — Pyrola asarifolia
  - **PYPI** — Pyrola picta
  - **RAUN2** — Ranunculus uncinatus
  - **RHMA** — Rhododendron macrophyllum
  - **RIBES** — Ribes
  - **RILO** — Ribes lobbii
  - **ROGY** — Rosa gymnocarpa
  - **RUBUS** — Rubus
  - **RULE** — Rubus leucodermis
  - **RUNI** — Rubus nivalis
  - **RUPA** — Rubus parviflorus
  - **RUUR** — Rubus ursinus
  - **SACE** — Sambucus cerulea
  - **SEJA** — Senecio jacobaea
  - **SESY** — Senecio sylvaticus
  - **SIAN2** — Silene antirrhina
  - **SOAR** — Sonchus arvensis
  - **STELL** — Stellaria
  - **STCA2** — Stellaria calycantha
  - **TABR** — Taxus brevifolia
  - **THPL** — Thuja plicata
  - **TITR** — Tiarella trifoliata
  - **TRLA2** — Trientalis latifolia
  - **TROV** — Trillium ovatum
  - **TRCA** — Trisetum canescens
  - **TRCE** — Trisetum cernuum
  - **TSHE** — Tsuga heterophylla
  - **VAME** — Vaccinium membranaceum
  - **VAPA** — Vaccinium parvifolium
  - **VAHE** — Vancouveria hexandra
  - **VEAR** — Veronica arvensis
  - **VESE** — Veronica serpyllifolia
  - **VISE** — Viola sempervirens
  - **WHMO** — Whipplea modesta
  - **CACH** — Castanopsis chrysophylla
  - **EBAU** — Eburophyton austiniae
  - **SADO** — Satureja douglasii
  - **SMST** — Smilacina stellata
  - **EPAN** — Epilobium angustifolium
  - **SYMO** — Symphoricarpos mollis
  - **GNMI** — Gnaphalium microcephalum
  - **LUCA2** — Luzula campestris
  - **UNKN** — unknown plant
  - **HERB** — Herb
  - **TREE** — Tree
  - **SHRUB** — Shrub
  - **MOSS** — Moss
  - **LICHEN** — Lichen
  - **GRASS** — Grass
  - **LITTER** — Litter
  - **CWD** — Coarse woody debris
  - **BARE** — Bare ground
  - **LOG** — Log
  - **STONE** — Stone
  - **BUTT** — Tree butt
  - **STUMP** — Tree stump
  - **SNAG** — Snag
  - **UNKN2** — unknown plant 2
  - **ROOT** — Root


- **EQUATION** — varchar, freetext

Explicit equation used for calculating biomass. mht = height, mcov= cover, mdba = bottom diameter


- **ALTEQU** — varchar, enum

Alternative equation within a species for calculating biomass

  - **1** — primary equation
  - **2** — second alternative equation
  - **3** — third alternative equation
  - **4** — fourth alternative equation


## Entity 6: Estimated biomass using measurement data and equations

### Attribute list

- **DBCODE** — varchar, enum

Database code

  - **TP103** — FSDB database code TP103


- **ENTITY** — numeric(1.000000), range  6.0000  6.0000, number

Entity number


- **STUDY** — varchar, enum

Study site location

  - **Starrbright** — Starrbright Timber Sale


- **YEAR** — numeric(1.000000), range  1992.0000  2024.0000, number

Sampling year


- **SEASON** — varchar, enum

Sampling season (1 - 3)

  - **U** — Unknown season of sampling
  - **1** — First season of sampling per year (usually june)
  - **2** — Second season of sampling per year (usually july)
  - **3** — Third season of sampling per year (usually august)


- **BLOCK** — numeric(1.000000), range  1.0000  25.0000, number

Sample block (1-25)


- **PLOT** — numeric(1.000000), range  1.0000  9.0000, number

Sample plot w/in block (1-9)


- **TMT** — varchar, enum

Treatment code (A-I)

  - **A** — Control
  - **B** — Senecio sylvaticus removed
  - **C** — Epilobium angustifolium removed
  - **D** — Senecio sylvaticus and Epilobium angustifolium removed
  - **E** — Senecio sylvaticus and community removed
  - **F** — Epilobium angustifolium and community removed
  - **G** — Community removed
  - **H** — Rubus ursinus removed
  - **I** — Berberis nervosa and Gaultheria shallon removed


- **SPECIES** — varchar, enum

Species acronym according to Garrison et al. 1974

  - **BORAGI** — Boraginaceae
  - **LILIAC** — Liliaceae
  - **ABGR** — Abies grandis
  - **ACCI** — Acer circinatum
  - **ACMA** — Acer macrophyllum
  - **ACTR** — Achlys triphylla
  - **ADBI** — Adenocaulon bicolor
  - **AGGR** — Agoseris grandiflora
  - **AGHE** — Agoseris heterophylla
  - **AGROS** — Agrostis
  - **AGEX** — Agrostis exarata
  - **AGTE** — Agrostis tenuis
  - **AICA** — Aira caryophyllea
  - **AMAL** — Amelanchier alnifolia
  - **ANMA** — Anaphalis margaritacea
  - **ANDE** — Anemone deltoidea
  - **ANLY2** — Anemone lyallii
  - **ARME** — Arbutus menziesii
  - **ARCO3** — Arctostaphylos columbiana
  - **AREL** — Arrhenatherum elatius
  - **ASCA3** — Asarum caudatum
  - **BENE** — Berberis nervosa
  - **BROMU** — Bromus
  - **BRVU** — Bromus vulgaris
  - **CASC2** — Campanula scouleri
  - **CARDA** — Cardamine
  - **CAOL** — Cardamine oligosperma
  - **CAAM** — Carex amplifolia
  - **CEANO** — Ceanothus
  - **CEIN** — Ceanothus integerrimus
  - **CESA** — Ceanothus sanguineus
  - **CEVE** — Ceanothus velutinus
  - **CERAS** — Cerastium
  - **CEVU** — Cerastium vulgatum
  - **CHME** — Chimaphila menziesii
  - **CHUM** — Chimaphila umbellata
  - **CHLE2** — Chrysanthemum leucanthemum
  - **CIRSI** — Cirsium
  - **CIAR** — Cirsium arvense
  - **CIVU** — Cirsium vulgare
  - **CLUN** — Clintonia uniflora
  - **COHE** — Collomia heterophylla
  - **COCA2** — Conyza canadensis
  - **CONU** — Cornus nuttallii
  - **CRCA** — Crepis capillaris
  - **DEEL** — Deschampsia elongata
  - **ELGL** — Elymus glaucus
  - **EPILO** — Epilobium
  - **EPMI** — Epilobium minutum
  - **EPPA** — Epilobium paniculatum
  - **EPWA** — Epilobium watsonii
  - **EQTE** — Equisetum telmateia
  - **FESTU** — Festuca
  - **FEMY** — Festuca myuros
  - **FEOC** — Festuca occidentalis
  - **FRVE** — Fragaria vesca
  - **GALIU** — Galium
  - **GATR** — Galium triflorum
  - **GAFR** — Garrya fremontii
  - **GASH** — Gaultheria shallon
  - **GNPU** — Gnaphalium purpureum
  - **GNUL** — Gnaphalium uliginosum
  - **GOOB** — Goodyera oblongifolia
  - **HIAL** — Hieracium albiflorum
  - **HOLA** — Holcus lanatus
  - **HODI** — Holodiscus discolor
  - **HYPE** — Hypericum perforatum
  - **HYRA** — Hypochaeris radicata
  - **LAMU** — Lactuca muralis
  - **LASE** — Lactuca serriola
  - **LIBO2** — Linnaea borealis
  - **LISTE** — Listera
  - **LICO3** — Listera cordata
  - **LOCI** — Lonicera ciliosa
  - **LOCR** — Lotus crassifolius
  - **LOMI** — Lotus micranthus
  - **LOPU** — Lotus purshianus
  - **LULA** — Lupinus latifolius
  - **MAGR** — Madia gracilis
  - **MOPE** — Montia perfoliata
  - **MOSI** — Montia sibirica
  - **MYDI** — Myosotis discolor
  - **NEPA** — Nemophila parviflora
  - **OSCH** — Osmorhiza chilensis
  - **PEFR2** — Petasites frigidus
  - **PILA** — Pinus lambertiana
  - **POLYG** — Polygonum
  - **PODO** — Polygonum douglasii
  - **POMU** — Polystichum munitum
  - **PRVU** — Prunella vulgaris
  - **PREM** — Prunus emarginata
  - **PSME** — Pseudotsuga menziesii
  - **PTAQ** — Pteridium aquilinum
  - **PYAS** — Pyrola asarifolia
  - **PYPI** — Pyrola picta
  - **RAUN2** — Ranunculus uncinatus
  - **RHMA** — Rhododendron macrophyllum
  - **RIBES** — Ribes
  - **RILO** — Ribes lobbii
  - **ROGY** — Rosa gymnocarpa
  - **RUBUS** — Rubus
  - **RULE** — Rubus leucodermis
  - **RUNI** — Rubus nivalis
  - **RUPA** — Rubus parviflorus
  - **RUUR** — Rubus ursinus
  - **SACE** — Sambucus cerulea
  - **SEJA** — Senecio jacobaea
  - **SESY** — Senecio sylvaticus
  - **SIAN2** — Silene antirrhina
  - **SOAR** — Sonchus arvensis
  - **STELL** — Stellaria
  - **STCA2** — Stellaria calycantha
  - **TABR** — Taxus brevifolia
  - **THPL** — Thuja plicata
  - **TITR** — Tiarella trifoliata
  - **TRLA2** — Trientalis latifolia
  - **TROV** — Trillium ovatum
  - **TRCA** — Trisetum canescens
  - **TRCE** — Trisetum cernuum
  - **TSHE** — Tsuga heterophylla
  - **VAME** — Vaccinium membranaceum
  - **VAPA** — Vaccinium parvifolium
  - **VAHE** — Vancouveria hexandra
  - **VEAR** — Veronica arvensis
  - **VESE** — Veronica serpyllifolia
  - **VISE** — Viola sempervirens
  - **WHMO** — Whipplea modesta
  - **CACH** — Castanopsis chrysophylla
  - **EBAU** — Eburophyton austiniae
  - **SADO** — Satureja douglasii
  - **SMST** — Smilacina stellata
  - **EPAN** — Epilobium angustifolium
  - **SYMO** — Symphoricarpos mollis
  - **GNMI** — Gnaphalium microcephalum
  - **LUCA2** — Luzula campestris
  - **UNKN** — unknown plant
  - **HERB** — Herb
  - **TREE** — Tree
  - **SHRUB** — Shrub
  - **MOSS** — Moss
  - **LICHEN** — Lichen
  - **GRASS** — Grass
  - **LITTER** — Litter
  - **CWD** — Coarse woody debris
  - **BARE** — Bare ground
  - **LOG** — Log
  - **STONE** — Stone
  - **BUTT** — Tree butt
  - **STUMP** — Tree stump
  - **SNAG** — Snag
  - **UNKN2** — unknown plant 2
  - **ROOT** — Root


- **B_VAL** — numeric(1), range  1.0000  8.0000, number

Unique biomass value for a given year, season, block, plot, tmt, species.


- **BIOMASS** — numeric(0.010000), range  0.0000  2279755.7500, grams per square meter

Estimated biomass


## Entity 7: Assessment of 2018 burn severity

### Attribute list

- **DBCODE** — varchar, enum

Database code

  - **TP103** — FSDB database code TP103


- **ENTITY** — numeric(1.000000), range  7.0000  7.0000, number

Entity number


- **STUDY** — varchar, enum

Study site location

  - **Starrbright** — Starrbright Timber Sale


- **YEAR** — numeric(1.000000), range  2019.0000  2019.0000, number

Sampling year


- **SDATE** — datetime, freetext

Vegetation sampling date


- **BLOCK** — numeric(1.000000), range  1.0000  25.0000, number

Sample block (1-25)


- **PLOT** — numeric(1.000000), range  1.0000  9.0000, number

Sample plot w/in block (1-9)


- **TMT** — varchar, enum

Treatment code (A-I)

  - **A** — Control
  - **B** — Senecio sylvaticus removed
  - **C** — Epilobium angustifolium removed
  - **D** — Senecio sylvaticus and Epilobium angustifolium removed
  - **E** — Senecio sylvaticus and community removed
  - **F** — Epilobium angustifolium and community removed
  - **G** — Community removed
  - **H** — Rubus ursinus removed
  - **I** — Berberis nervosa and Gaultheria shallon removed


- **TREES_LIVE** — numeric(1), range  0.0000  12.0000, number

Number of live trees within 2.5 m of the plot


- **TREES_DEAD** — numeric(1), range  0.0000  11.0000, number

Number of dead trees within 2.5 m of the plot


- **PROP_DEAD** — numeric(1), range  0.0000  100.0000, number

Proportion of trees within 2.5 m of the plot that are dead (undefined if there are no trees)


- **PCT_BURN** — numeric(1), range  0.0000  100.0000, percent

Percentage of ground surface burned or charred within the plot


- **PCT_BURN_ADJ** — numeric(1), range  0.0000  100.0000, percent

Percentage of ground surface burned or charred within 0.5 m of the plot edge


- **BURN_CLASS** — varchar, enum

Burn class

  - **B** — Burned
  - **PB** — Partially burned
  - **UB** — Unburned


- **BURN_COMMENT** — varchar, freetext

Description of burn conditions


## Attribute Definitions

**ALTEQU**

Alternative equation within a species for calculating biomass


**BIOMASS**

Estimated biomass


**BLOCK**

Sample block (1-25)


**BURN_CLASS**

Burn class


**BURN_COMMENT**

Description of burn conditions


**B_VAL**

Unique biomass value for a given year, season, block, plot, tmt, species.


**COV1_4TH**

Cover in the "quarter" plot (0.25 x 0.25 m), used to subsample when species density was high


**COVER**

Cover in the 1 m2 plot


**DBA**

Basal diameter


**DBCODE**

Database code


**DENSITYM**

Density of mature stems/sprouts (1 m2 plot)


**DENSITYS**

Density of seedlings (1 m2 plot)


**ENTITY**

Entity number


**EQUATION**

Explicit equation used for calculating biomass. mht = height, mcov= cover, mdba = bottom diameter


**HT**

Total height (including inflorescence if present)


**HTV**

Height of vegetative portion


**LC**

Line count


**NOTES**

Comments


**NREMBUF**

Number of stems removed from 75 cm buffer


**NREMPLT**

Number of stems removed from 1 m2 plot


**PCT_BURN**

Percentage of ground surface burned or charred within the plot


**PCT_BURN_ADJ**

Percentage of ground surface burned or charred within 0.5 m of the plot edge


**PLOT**

Sample plot w/in block (1-9)


**PROP_DEAD**

Proportion of trees within 2.5 m of the plot that are dead (undefined if there are no trees)


**RDATE**

Date of species removal (yyyymmdd)


**REC_NO**

Unique record number for a given year, season, block, plot, tmt, species and stratum


**SDATE**

Vegetation sampling date


**SEASON**

Sampling season (1 - 3)


**SPECIES**

Species acronym according to Garrison et al. 1974


**STEM1BV1**

Biomass parameter 1 of stem1


**STEM1BV2**

Biomass parameter 2 of stem1


**STEM1NO**

Number of stems containing the corresponding bv1 and bv2 values


**STEM2BV1**

Biomass parameter 1 of stem2


**STEM2BV2**

Biomass parameter 2 of stem2


**STEM2NO**

Number of stems containing the corresponding bv1 and bv2 values


**STEM3BV1**

Biomass parameter 1 of stem3


**STEM3BV2**

Biomass parameter 2 of stem3


**STEM3NO**

Number of stems containing the corresponding bv1 and bv2 values


**STEM4BV1**

Biomass parameter 1 of stem4


**STEM4BV2**

Biomass parameter 2 of stem4


**STEM5BV1**

Biomass parameter 1 of stem5


**STEM5BV2**

Biomass parameter 2 of stem5


**STRATUM**

Pre-harvest tree stratum: (O)verstory, (S)ubcanopy, (U)nderstory


**STUDY**

Study site location


**TMT**

Treatment code (A-I)


**TREES_DEAD**

Number of dead trees within 2.5 m of the plot


**TREES_LIVE**

Number of live trees within 2.5 m of the plot


**YEAR**

Sampling year


## Enumerated Domains

### DBCODE

- **TP103** — FSDB database code TP103

### STUDY

- **Starrbright** — Starrbright Timber Sale

### SEASON

- **U** — Unknown season of sampling

- **1** — First season of sampling per year (usually june)

- **2** — Second season of sampling per year (usually july)

- **3** — Third season of sampling per year (usually august)

### TMT

- **A** — Control

- **B** — Senecio sylvaticus removed

- **C** — Epilobium angustifolium removed

- **D** — Senecio sylvaticus and Epilobium angustifolium removed

- **E** — Senecio sylvaticus and community removed

- **F** — Epilobium angustifolium and community removed

- **G** — Community removed

- **H** — Rubus ursinus removed

- **I** — Berberis nervosa and Gaultheria shallon removed

### SPECIES

- **BORAGI** — Boraginaceae

- **LILIAC** — Liliaceae

- **ABGR** — Abies grandis

- **ACCI** — Acer circinatum

- **ACMA** — Acer macrophyllum

- **ACTR** — Achlys triphylla

- **ADBI** — Adenocaulon bicolor

- **AGGR** — Agoseris grandiflora

- **AGHE** — Agoseris heterophylla

- **AGROS** — Agrostis

- **AGEX** — Agrostis exarata

- **AGTE** — Agrostis tenuis

- **AICA** — Aira caryophyllea

- **AMAL** — Amelanchier alnifolia

- **ANMA** — Anaphalis margaritacea

- **ANDE** — Anemone deltoidea

- **ANLY2** — Anemone lyallii

- **ARME** — Arbutus menziesii

- **ARCO3** — Arctostaphylos columbiana

- **AREL** — Arrhenatherum elatius

- **ASCA3** — Asarum caudatum

- **BENE** — Berberis nervosa

- **BROMU** — Bromus

- **BRVU** — Bromus vulgaris

- **CASC2** — Campanula scouleri

- **CARDA** — Cardamine

- **CAOL** — Cardamine oligosperma

- **CAAM** — Carex amplifolia

- **CEANO** — Ceanothus

- **CEIN** — Ceanothus integerrimus

- **CESA** — Ceanothus sanguineus

- **CEVE** — Ceanothus velutinus

- **CERAS** — Cerastium

- **CEVU** — Cerastium vulgatum

- **CHME** — Chimaphila menziesii

- **CHUM** — Chimaphila umbellata

- **CHLE2** — Chrysanthemum leucanthemum

- **CIRSI** — Cirsium

- **CIAR** — Cirsium arvense

- **CIVU** — Cirsium vulgare

- **CLUN** — Clintonia uniflora

- **COHE** — Collomia heterophylla

- **COCA2** — Conyza canadensis

- **CONU** — Cornus nuttallii

- **CRCA** — Crepis capillaris

- **DEEL** — Deschampsia elongata

- **ELGL** — Elymus glaucus

- **EPILO** — Epilobium

- **EPMI** — Epilobium minutum

- **EPPA** — Epilobium paniculatum

- **EPWA** — Epilobium watsonii

- **EQTE** — Equisetum telmateia

- **FESTU** — Festuca

- **FEMY** — Festuca myuros

- **FEOC** — Festuca occidentalis

- **FRVE** — Fragaria vesca

- **GALIU** — Galium

- **GATR** — Galium triflorum

- **GAFR** — Garrya fremontii

- **GASH** — Gaultheria shallon

- **GNPU** — Gnaphalium purpureum

- **GNUL** — Gnaphalium uliginosum

- **GOOB** — Goodyera oblongifolia

- **HIAL** — Hieracium albiflorum

- **HOLA** — Holcus lanatus

- **HODI** — Holodiscus discolor

- **HYPE** — Hypericum perforatum

- **HYRA** — Hypochaeris radicata

- **LAMU** — Lactuca muralis

- **LASE** — Lactuca serriola

- **LIBO2** — Linnaea borealis

- **LISTE** — Listera

- **LICO3** — Listera cordata

- **LOCI** — Lonicera ciliosa

- **LOCR** — Lotus crassifolius

- **LOMI** — Lotus micranthus

- **LOPU** — Lotus purshianus

- **LULA** — Lupinus latifolius

- **MAGR** — Madia gracilis

- **MOPE** — Montia perfoliata

- **MOSI** — Montia sibirica

- **MYDI** — Myosotis discolor

- **NEPA** — Nemophila parviflora

- **OSCH** — Osmorhiza chilensis

- **PEFR2** — Petasites frigidus

- **PILA** — Pinus lambertiana

- **POLYG** — Polygonum

- **PODO** — Polygonum douglasii

- **POMU** — Polystichum munitum

- **PRVU** — Prunella vulgaris

- **PREM** — Prunus emarginata

- **PSME** — Pseudotsuga menziesii

- **PTAQ** — Pteridium aquilinum

- **PYAS** — Pyrola asarifolia

- **PYPI** — Pyrola picta

- **RAUN2** — Ranunculus uncinatus

- **RHMA** — Rhododendron macrophyllum

- **RIBES** — Ribes

- **RILO** — Ribes lobbii

- **ROGY** — Rosa gymnocarpa

- **RUBUS** — Rubus

- **RULE** — Rubus leucodermis

- **RUNI** — Rubus nivalis

- **RUPA** — Rubus parviflorus

- **RUUR** — Rubus ursinus

- **SACE** — Sambucus cerulea

- **SEJA** — Senecio jacobaea

- **SESY** — Senecio sylvaticus

- **SIAN2** — Silene antirrhina

- **SOAR** — Sonchus arvensis

- **STELL** — Stellaria

- **STCA2** — Stellaria calycantha

- **TABR** — Taxus brevifolia

- **THPL** — Thuja plicata

- **TITR** — Tiarella trifoliata

- **TRLA2** — Trientalis latifolia

- **TROV** — Trillium ovatum

- **TRCA** — Trisetum canescens

- **TRCE** — Trisetum cernuum

- **TSHE** — Tsuga heterophylla

- **VAME** — Vaccinium membranaceum

- **VAPA** — Vaccinium parvifolium

- **VAHE** — Vancouveria hexandra

- **VEAR** — Veronica arvensis

- **VESE** — Veronica serpyllifolia

- **VISE** — Viola sempervirens

- **WHMO** — Whipplea modesta

- **CACH** — Castanopsis chrysophylla

- **EBAU** — Eburophyton austiniae

- **SADO** — Satureja douglasii

- **SMST** — Smilacina stellata

- **EPAN** — Epilobium angustifolium

- **SYMO** — Symphoricarpos mollis

- **GNMI** — Gnaphalium microcephalum

- **LUCA2** — Luzula campestris

- **UNKN** — unknown plant

- **HERB** — Herb

- **TREE** — Tree

- **SHRUB** — Shrub

- **MOSS** — Moss

- **LICHEN** — Lichen

- **GRASS** — Grass

- **LITTER** — Litter

- **CWD** — Coarse woody debris

- **BARE** — Bare ground

- **LOG** — Log

- **STONE** — Stone

- **BUTT** — Tree butt

- **STUMP** — Tree stump

- **SNAG** — Snag

- **UNKN2** — unknown plant 2

- **ROOT** — Root

### STRATUM

- **O** — Overstory layer

- **S** — Subcanopy layer

- **U** — Understory layer

- **N** — Not applicable; STRATUM applies only to tree species in the pre-harvest sample (1990)

### ALTEQU

- **1** — primary equation

- **2** — second alternative equation

- **3** — third alternative equation

- **4** — fourth alternative equation

### BURN_CLASS

- **B** — Burned

- **PB** — Partially burned

- **UB** — Unburned