SA029: Western columbine genetics across HJ Andrews Experimental Forest meadow communities

Citation

Jones, A.; Gannon, D. 2019. Western columbine genetics across HJ Andrews Experimental Forest meadow communities Long-Term Ecological Research Andrews Forest LTER Site. [Database]. Available: https://andrewsforest-stage.forestry.oregonstate.edu/data/fsdb-data-catalog/SA029 Accessed 2026-05-10.

Abstract

Woody plant encroachment is diminishing meadow and grassland habitat on a global scale. Increased woody cover influences local conditions such as light/shade environments, local soil characteristics, understory plant community structure, and disturbance regimes. Woody encroachment may also affect landscape-scale biological processes, such as herbaceous plant population structure, through reducing the total cover and continuity of open habitat and eroding mutualistic interactions, such as plant-pollinator interaction networks. A major concern is that habitat fragmentation will have a cascading effect if one or more mutualistic partners is adversely affected. For example, if pollinators are sensitive to disturbance, fragmentation may reduce rates of gene flow among sub-populations of plants, which is predicted to decrease effective population sizes and diminish adaptive potential (i.e. the capacity to respond to selective pressures through the evolution of genetically-based and heritable traits). Alpine meadows of the Cascade Mountains, which support diverse wildflower and pollinator communities, have shrunk dramatically over the last century as a result of forest encroachment. We posited that, as meadows become smaller and less connected, pollinators may abandon the smallest meadows and focus foraging efforts on the largest, most connected meadows with the most resources. This could expedite the decline and ultimate collapse of meadow communities through reducing adaptive potential across sub-populations of plants. We focus on a plant-pollinator interaction between a common, nectar-producing plant, Aquilegia formosa (western columbine), and rufous hummingbird (Selasphorus rufus) pollinators in four montane meadow complexes in the H.J. Andrews experimental forest, Oregon, USA (HJA). Using hummingbird movement data from SA028 (see H.J. Andrews project database), we first ask whether further forest encroachment in the HJA may alter hummingbird movement patterns among meadows. Secondly, using genotyping-by-sequencing to query the genomes of A. formosa individuals from 28 meadows representing a gradient of small and isolated meadows to large, connected meadows, we ask whether contemporary hummingbird movements predict plant population genetic structure and connectivity. A strong hummingbird preference for open habitat and support for a model predicting genetic distances among plants with hummingbird movement over standard geographic distances would support our hypothesis. Alternative results may suggest more robust plant-pollinator interactions in the face of woody encroachment. Western columbine leaves were collected from 28 meadows within the H. J. Andrews Experimental Forest, Oregon, USA. We sampled six meadows on Lookout Mountain, twelve along Frissell Ridge, six on Carpenter Mountain, and two clearings along forest road 506.

Coverage

Temporal coverage: 2019-06-21 to 2019-06-26

Geographic coverage: Montane meadows within the H. J. Andrews Experimental Forest, Western Cascades, Oregon, USA.

Spatial coverage:

Loading map…

Bounds: W -122.15197140, E -122.10287200, N 44.20325610, S 44.28345743

Purpose

Assess the degree to which woody encroachment on montane meadow communities may disrupt plant-pollinator interactions and influence meadow plant population connectivity and viability.

Project

Title: Long-Term Ecological Research

Personnel

Posy Elizabeth Busby - Principal Investigator
Assistant Professor OSU Botany & Plant Pathology

Email: busbyp@science.oregonstate.edu, posybusby@gmail.com

URL: http://bpp.oregonstate.edu/content/posy-busby

ORCID: http://orcid.org/0000-0002-2837-9820
Matthew G Betts - Principal Investigator
Department of Forest Ecosystems and Society; 201E Richardson Hall; College of Forestry; Oregon State University, Corvallis, OR, 97331

Phone: (541) 737-3841

Email: matt.betts@oregonstate.edu

URL: http://www.fsl.orst.edu/flel/index.htm

ORCID: http://orcid.org/0000-0002-7100-2551
Brooke E. Penaluna - Principal Investigator
Email: brooke.penaluna@usda.gov, Brooke.Penaluna@oregonstate.edu

URL: https://www.fs.fed.us/pnw/lwm/aem/people/penaluna.html

ORCID: http://orcid.org/0000-0001-7215-770X
Catalina Segura - Principal Investigator
Assistant Professor; Department of Forest Engineering, Resources, and Management; Oregon State University, Corvallis, OR, 97331

Phone: 541-737-6568

Email: catalina.segura@oregonstate.edu

URL: http://ferm.forestry.oregonstate.edu/facstaff/segura-catalina

ORCID: http://orcid.org/0000-0002-0924-1172
David Bell - Principal Investigator
Email: david.bell@usda.gov, david.bell@oregonstate.edu

URL: https://lemma.forestry.oregonstate.edu/about/david-bell

ORCID: http://orcid.org/0000-0002-2673-5836

Abstract

The H.J. Andrews Experimental Forest is a living laboratory that provides unparalleled opportunities for the study of forest and stream ecosystems in the central Cascade Range of Oregon. Since 1980, as a part of the National Science Foundation Long Term Ecological Research (NSF-LTER) program, the Andrews Experimental Forest has become a leader in the analysis of forest and stream ecosystem dynamics.
Long-term field experiments and measurement programs have focused on climate dynamics, streamflow, water quality, and vegetation succession. Currently researchers are working to develop concepts and tools needed to predict effects of natural disturbance, land use, and climate change on ecosystem structure, function, and species composition.
The Andrews Experimental Forest is administered cooperatively by the USDA Forest Service Pacific Northwest Research Station, Oregon State University and the Willamette National Forest. Funding for the research program comes from the National Science Foundation (NSF), US Forest Service Pacific Northwest Research Station, Oregon State University, and other sources.

Funding

Data were provided by the HJ Andrews Experimental Forest research program, funded by the National Science Foundation's Long-Term Ecological Research Program (DEB 2025755), US Forest Service Pacific Northwest Research Station, and Oregon State University. National Science Foundation: DEB2025755

Study Area Description

Long-Term Ecological Research
The Andrews Forest is situated in the western Cascade Range of Oregon, and covers the entire 15,800-acre (6400-ha) drainage basin of Lookout Creek. Elevation ranges from 1350 to 5340 feet (410 to 1630 m). Broadly representative of the rugged mountainous landscape of the Pacific Northwest, the Andrews Forest contains excellent examples of the region's conifer forests and associated wildlife and stream ecosystems. These forests are among the tallest and most productive in the world, with tree heights of often greater than 250 ft (75 m). Streams are steep, cold and clean, providing habitat for numerous aquatic organisms.

Associated Party

Andy Jones
Role: Principal Investigator

2070 Cordley Hall, Corvallis, Oregon, 97330, USA

Email: jonesfr@science.oregonstate.edu

URL: http://blogs.oregonstate.edu/joneslab/
Matthew G Betts
Role: Other Researcher

Department of Forest Ecosystems and Society; 201E Richardson Hall; College of Forestry; Oregon State University, Corvallis, OR, 97331

Phone: (541) 737-3841

Email: matt.betts@oregonstate.edu

URL: http://www.fsl.orst.edu/flel/index.htm
Andy Jones
Role: Creator

2070 Cordley Hall, Corvallis, Oregon, 97330, USA

Email: jonesfr@science.oregonstate.edu

URL: http://blogs.oregonstate.edu/joneslab/
Dustin Gannon
Role: Creator

Email: dustygannon93@gmail.com

Contact

Information Manager
Andrews Forest LTER Program, US Forest Service Pacific Northwest Research Station, 3200 SW Jefferson Way, Corvallis, OR, 97331

Email: hjaweb@lists.oregonstate.edu

URL: http://andrewsforest.oregonstate.edu/
Dustin Gannon
Email: dustygannon93@gmail.com

Publisher

Andrews Forest LTER Site
Role: Publisher

Forest Ecosystems and Society Department in Forestry, Oregon State University, 201K Richardson Hall, Corvallis, OR, 97331-5752

Phone: (541) 737-8480

Email: lterweb@lists.oregonstate.edu

URL: http://andrewsforest.oregonstate.edu/

Study Description

Woody plant encroachment is diminishing meadow and grassland habitat on a global scale. Increased woody cover influences local conditions such as light/shade environments, local soil characteristics, understory plant community structure, and disturbance regimes. Woody encroachment may also affect landscape-scale biological processes, such as herbaceous plant population structure, through reducing the total cover and continuity of open habitat and eroding mutualistic interactions, such as plant-pollinator interaction networks. A major concern is that habitat fragmentation will have a cascading effect if one or more mutualistic partners is adversely affected. For example, if pollinators are sensitive to disturbance, fragmentation may reduce rates of gene flow among sub-populations of plants, which is predicted to decrease effective population sizes and diminish adaptive potential (i.e. the capacity to respond to selective pressures through the evolution of genetically-based and heritable traits). Alpine meadows of the Cascade Mountains, which support diverse wildflower and pollinator communities, have shrunk dramatically over the last century as a result of forest encroachment. We posited that, as meadows become smaller and less connected, pollinators may abandon the smallest meadows and focus foraging efforts on the largest, most connected meadows with the most resources. This could expedite the decline and ultimate collapse of meadow communities through reducing adaptive potential across sub-populations of plants. We focus on a plant-pollinator interaction between a common, nectar-producing plant, Aquilegia formosa (western columbine), and rufous hummingbird (Selasphorus rufus) pollinators in four montane meadow complexes in the H.J. Andrews experimental forest, Oregon, USA (HJA). Using hummingbird movement data from SA028 (see H.J. Andrews project database), we first ask whether further forest encroachment in the HJA may alter hummingbird movement patterns among meadows. Secondly, using genotyping-by-sequencing to query the genomes of A. formosa individuals from 28 meadows representing a gradient of small and isolated meadows to large, connected meadows, we ask whether contemporary hummingbird movements predict plant population genetic structure and connectivity. A strong hummingbird preference for open habitat and support for a model predicting genetic distances among plants with hummingbird movement over standard geographic distances would support our hypothesis. Alternative results may suggest more robust plant-pollinator interactions in the face of woody encroachment. Western columbine leaves were collected from 28 meadows within the H. J. Andrews Experimental Forest, Oregon, USA. We sampled six meadows on Lookout Mountain, twelve along Frissell Ridge, six on Carpenter Mountain, and two clearings along forest road 506. Assess the degree to which woody encroachment on montane meadow communities may disrupt plant-pollinator interactions and influence meadow plant population connectivity and viability. Field Methods - SA029

Purpose: Assess the degree to which woody encroachment on montane meadow communities may disrupt plant-pollinator interactions and influence meadow plant population connectivity and viability.

Methods

Method Steps

Field Methods - SA029

In many cases, A. formosa did not occupy randomly selected meadows. In such cases, we visited the next meadow belonging to the same stratum and meadow complex in a randomized list. Due to time constraints, we were unable to sample all complex-stratum combinations. The final list of focal meadows included 28 meadows, with at least one meadow from each stratum in each complex. From the 28 meadows, we sampled fresh leaves from a minimum of six A. formosa individuals, placed them in 4ml cryotubes, and stored them in a charged Worthington CXR500 dry-shipper (-1900C) until transport back to Oregon State University where they could be stored in a -800C freezer.

Data Entry - SA029

Sample data (sample location, ID, meadow, meadow complex, forest cover, and phenological stage) were entered by hand. Entries were checked and verified in a separate data entry session.

Sampling

Study Extent

Sampling frequency: One-time leaf sample collection

Sampling Description

We sampled western columbine leaves (Aquilegia formosa) from four major meadow complexes (spatial clusters of meadows) within the H.J. Andrews Experimental Forest, Oregon, USA: two complexes along Frissel Ridge (M1 and M2), one on Carpenter Mountain (CM), and one on Lookout Mountain (LOM). Meadows within complexes were selected based on a two-step approach. First, we focused on meadows from which we had hummingbird RFID data (see project SA028) and those from which we had A. formosa seed set and/or pollen flow data. Because we aim to test the degree to which contemporary hummingbird movement predicts plant population genetic structure, we sought to sample plants in locations that should best reflect movement estimates. Divergence between the two is therefore less likely to be caused simply by sampling different meadows with different conditions, and more likely related to the biology of gene flow among plants on the landscape.
We then calculated meadow area (m2) and connectivity based on the amount of forest in a 100m-radius buffer around the meadow centroid (meadows with high proportions of forest cells surrounding them were assumed less connected). We stratified meadows into four categories: large-connected, large-isolated, small-connected, and small-isolated meadows based on natural but arbitrary breaks in the distributions of meadow size and connectivity (breaks were placed at 3.2 log10(m2) and 63% forest cover for meadow size and connectivity, respectively). Meadows from which we had complementary data (RFID or seed set data) were categorized into respective strata. We then filled the list in by randomly sampling within strata up to 32 meadows such that the final list included at least two meadows belonging to each stratum (large-connected, large-isolated, small-connected, small-isolated) within each of the four meadow complexes (M1, M2, CM, LOM).

Spatial Sampling Units

Software

No software entries listed in this EML file.

Keywords

LTER controlled vocabulary: genetics (theme), meadows (theme)

Taxonomic Hierarchy

All Organisms: All Organisms
Highest common category (ca. kingdom): Plantae
Division or Phylum: Magnoliophyta
Class: Magnoliopsida
Subclass: Magnoliidae
Order: Ranunculales
Family: Ranunculaceae
Genus: Aquilegia
Species: Aquilegia formosa

Data Entities

#	Entity	Metadata	Data
1	SA02901 Meadow characteristics Quantification of meadow area, proportion of forest, and stratum on meadow plots	METADATA	DATA
2	SA02902 Individual plant information and cover species cover, number of conspecifics, flower status, coordinates, and elevation of individual plant	METADATA	DATA

Metadata

SA02901 - Meadow characteristics

Object name: SA02901_v1.csv

Records: 28

Attributes: 10

Temporal coverage: 2019-06-21 to 2019-06-26

File size: 1716 byte

Checksum (MD5): 33ccb753843ab39ea29b0dc035397372

Format: headers=1, recordDelimiter=\r\n, fieldDelimiter=,, quoteCharacter=", orientation=column

Constraints (2)

primaryKey: PRIMARY
SA02901.COMPLEX, SA02901.MEADOW, SA02901.MEADOW_PLOT
notNullConstraint: NOTNULL
SA02901.COMPLEX, SA02901.DBCODE, SA02901.ENTITY, SA02901.MEADOW, SA02901.MEADOW_AREA, SA02901.MEADOW_PLOT, SA02901.PROP_FOREST100, SA02901.PROP_FOREST50, SA02901.PROP_FOREST500, SA02901.STRATUM

Attributes (10)

DBCODE - char(5) (nominal)

ID: SA02901.DBCODE

FSDB Database Code

Type system: Microsoft SQL Server 2019

Code definitions (1)

SA029
FSDB Database Study Code

ENTITY - numeric(2,0) (ratio)

ID: SA02901.ENTITY

Entity number

Type system: Microsoft SQL Server 2019

Unit: number

Precision: 1

Numeric domain: type=natural, min=1.0000 (exclusive=false), max=1.0000 (exclusive=false)

COMPLEX - char(10) (nominal)

ID: SA02901.COMPLEX

Cluster of meadows

Type system: Microsoft SQL Server 2019

Code definitions (3)

Carpenter
Carpenter Mountain Meadow Complex
Frissell
Frissell Roswell Ridge Meadow Complex
Lookout
Lookout Mountain Meadow Complex

MEADOW - char(3) (nominal)

ID: SA02901.MEADOW

Name of the meadow according to HJA meadows shape file

Type system: Microsoft SQL Server 2019

Code definitions (4)

CM
Carpenter Main Meadow
M2
Frissell Roswell Main 2 Meadow
LOM
Lookout Main Meadow
M1
Frissell Roswell Main 1 Meadow

MEADOW_PLOT - char(8) (nominal)

ID: SA02901.MEADOW_PLOT

meadow id for sampled area within a meadow (sub meadow)

Type system: Microsoft SQL Server 2019

Code definitions (28)

CMM2
Carpenter Main Meadow, plot m2
CMX11B
Carpenter Main Meadow, plot x11b
CMX11C
Carpenter Main Meadow, plot x11c
CMX41A
Carpenter Main Meadow, plot x41a
CMX41C
Carpenter Main Meadow, plot x41c
CMX43
Carpenter Main Meadow, plot x43
LOMCAGE
Lookout Main Meadow, plot m27
LOMLG1
Lookout Main Meadow, plot x15e
LOMM14
Lookout Main Meadow, plot x4a
LOMM27
Lookout Main Meadow, plot cage
LOMMWS
Lookout Main Meadow, plot lg1
LOMW1
Lookout Main Meadow, plot m14
LOMX15E
Lookout Main Meadow, plot w1
LOMX42
Lookout Main Meadow, plot ws
LOMX4A
Lookout Main Meadow, plot x42
M1LG1
Frissell Roswell Main 1 Meadow, plot lg1
M1M3
Frissell Roswell Main 1 Meadow, plot m3
M1M5
Frissell Roswell Main 1 Meadow, plot m5
M1X1A
Frissell Roswell Main 1 Meadow, plot x1a
M1X2B
Frissell Roswell Main 1 Meadow, plot x2b
M1X7
Frissell Roswell Main 1 Meadow, plot x7
M2M1
Frissell Roswell Main 2 Meadow, plot m1
M2X17
Frissell Roswell Main 2 Meadow, plot x17
M2X21A
Frissell Roswell Main 2 Meadow, plot x21a
M2X21B
Frissell Roswell Main 2 Meadow, plot x21b
M2X21E
Frissell Roswell Main 2 Meadow, plot x21e
M2X21F
Frissell Roswell Main 2 Meadow, plot x21f
M2X48
Frissell Roswell Main 2 Meadow, plot x48

MEADOW_AREA - numeric(7,1) (ratio)

ID: SA02901.MEADOW_AREA

Area covered by the meadow plot

Type system: Microsoft SQL Server 2019

Unit: square meters

Precision: 1

Numeric domain: type=real, min=128.0000 (exclusive=false), max=50000.0000 (exclusive=false)

PROP_FOREST50 - numeric(4,2) (ratio)

ID: SA02901.PROP_FOREST50

Proportion of forest within a 50 meter radius around the centroid of a meadow plot

Type system: Microsoft SQL Server 2019

Unit: number

Precision: 1

Numeric domain: type=real, min=0.0000 (exclusive=false), max=1.0000 (exclusive=false)

PROP_FOREST100 - numeric(4,2) (ratio)

ID: SA02901.PROP_FOREST100

Proportion of forest within a 100 meter radius around the centroid of a meadow plot

Type system: Microsoft SQL Server 2019

Unit: number

Precision: 1

Numeric domain: type=real, min=0.0000 (exclusive=false), max=1.0000 (exclusive=false)

PROP_FOREST500 - numeric(4,2) (ratio)

ID: SA02901.PROP_FOREST500

Proportion of forest within a 500 meter radius around the centroid of a meadow plot

Type system: Microsoft SQL Server 2019

Unit: number

Precision: 1

Numeric domain: type=real, min=0.0000 (exclusive=false), max=1.0000 (exclusive=false)

STRATUM - char(3) (nominal)

ID: SA02901.STRATUM

Stratum of the meadow plot

Type system: Microsoft SQL Server 2019

Code definitions (4)

LgC
large connected meadow
SmC
small connected meadow
LgI
large isolated meadow
SmI
small isolated meadow

SA02902 - Individual plant information and cover

Object name: SA02902_v1.csv

Records: 260

Attributes: 15

Temporal coverage: 2019-06-21 to 2019-06-26

File size: 21655 byte

Checksum (MD5): 132f02f454b821830ecbaa37f73e87ed

Format: headers=1, recordDelimiter=\r\n, fieldDelimiter=,, quoteCharacter=", orientation=column

Constraints (2)

primaryKey: PRIMARY
SA02902.COMPLEX, SA02902.MEADOW, SA02902.MEADOW_PLOT, SA02902.PLANT_ID, SA02902.SPECIES_CODE
notNullConstraint: NOTNULL
SA02902.COMPLEX, SA02902.COVER, SA02902.DBCODE, SA02902.ENTITY, SA02902.FLW_STATUS, SA02902.MEADOW, SA02902.MEADOW_PLOT, SA02902.PLANT_ID, SA02902.PL_IN_5M, SA02902.SAMPLEDATE, SA02902.SPECIES_CODE, SA02902.X_COORD, SA02902.Y_COORD

Attributes (15)

DBCODE - char(5) (nominal)

ID: SA02902.DBCODE

FSDB Database Code

Type system: Microsoft SQL Server 2019

Code definitions (1)

SA029
FSDB Database Study Code

ENTITY - numeric(2,0) (ratio)

ID: SA02902.ENTITY

Entity number

Type system: Microsoft SQL Server 2019

Unit: number

Precision: 1

Numeric domain: type=natural, min=2.0000 (exclusive=false), max=2.0000 (exclusive=false)

COMPLEX - char(10) (nominal)

ID: SA02902.COMPLEX

Cluster of meadows

Type system: Microsoft SQL Server 2019

Code definitions (3)

Carpenter
Carpenter Mountain Meadow Complex
Frissell
Frissell Roswell Ridge Meadow Complex
Lookout
Lookout Mountain Meadow Complex

MEADOW - char(3) (nominal)

ID: SA02902.MEADOW

Name of the meadow according to HJA meadows shape file

Type system: Microsoft SQL Server 2019

Code definitions (4)

CM
Carpenter Main Meadow
M2
Frissell Roswell Main 2 Meadow
LOM
Lookout Main Meadow
M1
Frissell Roswell Main 1 Meadow

MEADOW_PLOT - char(8) (nominal)

ID: SA02902.MEADOW_PLOT

meadow id for sampled area within a meadow (sub meadow)

Type system: Microsoft SQL Server 2019

Code definitions (28)

CMM2
Carpenter Main Meadow, plot m2
CMX11B
Carpenter Main Meadow, plot x11b
CMX11C
Carpenter Main Meadow, plot x11c
CMX41A
Carpenter Main Meadow, plot x41a
CMX41C
Carpenter Main Meadow, plot x41c
CMX43
Carpenter Main Meadow, plot x43
LOMCAGE
Lookout Main Meadow, plot m27
LOMLG1
Lookout Main Meadow, plot x15e
LOMM14
Lookout Main Meadow, plot x4a
LOMM27
Lookout Main Meadow, plot cage
LOMMWS
Lookout Main Meadow, plot lg1
LOMW1
Lookout Main Meadow, plot m14
LOMX15E
Lookout Main Meadow, plot w1
LOMX42
Lookout Main Meadow, plot ws
LOMX4A
Lookout Main Meadow, plot x42
M1LG1
Frissell Roswell Main 1 Meadow, plot lg1
M1M3
Frissell Roswell Main 1 Meadow, plot m3
M1M5
Frissell Roswell Main 1 Meadow, plot m5
M1X1A
Frissell Roswell Main 1 Meadow, plot x1a
M1X2B
Frissell Roswell Main 1 Meadow, plot x2b
M1X7
Frissell Roswell Main 1 Meadow, plot x7
M2M1
Frissell Roswell Main 2 Meadow, plot m1
M2X17
Frissell Roswell Main 2 Meadow, plot x17
M2X21A
Frissell Roswell Main 2 Meadow, plot x21a
M2X21B
Frissell Roswell Main 2 Meadow, plot x21b
M2X21E
Frissell Roswell Main 2 Meadow, plot x21e
M2X21F
Frissell Roswell Main 2 Meadow, plot x21f
M2X48
Frissell Roswell Main 2 Meadow, plot x48

SAMPLEDATE - datetime (dateTime)

ID: SA02902.SAMPLEDATE

Date of sampling

Type system: Microsoft SQL Server 2019

Date format: YYYY-MM-DD

SPECIES_CODE - char(4) (ordinal)

ID: SA02902.SPECIES_CODE

Code representing species scientific name

Type system: Microsoft SQL Server 2019

Code definitions (1)

AQFO
Aquilegia formosa

PLANT_ID - numeric(3,0) (ratio)

ID: SA02902.PLANT_ID

Unique plant ID for individual

Type system: Microsoft SQL Server 2019

Unit: number

Precision: 1

Numeric domain: type=natural, min=193.0000 (exclusive=false), max=479.0000 (exclusive=false)

X_COORD - numeric(6,0) (ratio)

ID: SA02902.X_COORD

X coordinate, WGS84, UTM Zone 10

Type system: Microsoft SQL Server 2019

Unit: number

Precision: 1

Numeric domain: type=natural, min=560406.0000 (exclusive=false), max=571679.0000 (exclusive=false)

Y_COORD - numeric(7,0) (ratio)

ID: SA02902.Y_COORD

Y coordinate WGS84, UTM Zone 10

Type system: Microsoft SQL Server 2019

Unit: number

Precision: 1

Numeric domain: type=natural, min=4893946.0000 (exclusive=false), max=4908747.0000 (exclusive=false)

ELEVATION - numeric(4,0) (ratio)

ID: SA02902.ELEVATION

meters above sea level

Type system: Microsoft SQL Server 2019

Unit: meters

Precision: 1

Numeric domain: type=whole, min=1268.0000 (exclusive=false), max=1545.0000 (exclusive=false)

PL_IN_5M - numeric(2,0) (ratio)

ID: SA02902.PL_IN_5M

number of conspecifics in a 5m radius from the focal plant

Type system: Microsoft SQL Server 2019

Unit: number

Precision: 1

Numeric domain: type=whole, min=0.0000 (exclusive=false), max=25.0000 (exclusive=false)

COVER - numeric(2,0) (ratio)

ID: SA02902.COVER

Overstory cover

Type system: Microsoft SQL Server 2019

Unit: percent

Precision: 1

Numeric domain: type=real, min=0.0000 (exclusive=false), max=100.0000 (exclusive=false)

FLW_STATUS - char(1) (nominal)

ID: SA02902.FLW_STATUS

Whether or not the plant had flowers at the time of sampling

Type system: Microsoft SQL Server 2019

Code definitions (3)

F
Flower buds, blooming flowers, or seed pods are present
V
Only vegetative structures present
N
Not noted

NOTES - varchar(25) (nominal)

ID: SA02902.NOTES

sample notes

Type system: Microsoft SQL Server 2019

Units


square meters	m2	area	meterSquared	meterSquared	1	square meters
percent	%	dimensionless	number	dimensionless	100	percent; a number
meters	m	length	meter	meter	1	meter; SI unit of length
number	number	dimensionless	number	dimensionless	1	dimensionless number, i.e., ratio, count

Related Datasets

SA028

Related Publications

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Related Files

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Intellectual Rights

Data Use Agreement:

The re-use of scientific data has the potential to greatly increase communication, collaboration and synthesis within and among disciplines, and thus is fostered, supported and encouraged. This Data Set is released under the Creative Commons license CC BY "Attribution" (see: https://creativecommons.org/licenses/by/4.0/). Creative Commons license CC BY - Attribution is a license that allows others to distribute, remix, tweak, and build upon your work (even commercially), as long as you are credited for the original creation. This license accommodates maximum dissemination and use of licensed materials.

It is considered professional conduct and an ethical obligation to acknowledge the work of other scientists. The Data User is asked to provide attribution of the original work if this data package is shared in whole or by individual parts or used in the derivation of other products. A recommended citation is provided for each Data Set in the Andrews LTER data catalog (see: http://andlter.forestry.oregonstate.edu/data/catalog/datacatalog.aspx). A generic citation is also provided for this Data Set on the website https://portal.edirepository.org in the summary metadata page. Data Users are thus strongly encouraged to consider consultation, collaboration and/or co-authorship with the Data Set Creator.

While substantial efforts are made to ensure the accuracy of data and associated documentation, complete accuracy of data sets cannot be guaranteed and all data are made available "as is." The Data User should be aware, however, that data are updated periodically and it is the responsibility of the Data User to check for new versions of the data. The data authors and the repository where these data were obtained shall not be liable for damages resulting from any use or misinterpretation of the data.

General acknowledgement: Data were provided by the HJ Andrews Experimental Forest research program, funded by the National Science Foundation's Long-Term Ecological Research Program (DEB 2025755), US Forest Service Pacific Northwest Research Station, and Oregon State University.

Licensed

License: N/A

Maintenance

Maintenance update frequency: notPlanned

Description

An update history is logged and maintained with each new version of every dataset.

Change History

Version1 (2017-05-19)
Study code and preliminary metadata established
Version2 (2019-12-31)
Uploaded new data to SQL and prepared for PASTA. Originally uploaded to PASTA before this update record was added. This is fixing last update date.

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SA029: Western columbine genetics across HJ Andrews Experimental Forest meadow communities

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